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Habit Herbs annual, glandular puberulent and sparsely villous. Herbs annual, densely villous and strigillose, glandular puberulent on distal parts.
Stems

slender, unbranched or branched, 10–20 cm.

well branched, 15–50 cm.

Leaves

primarily cauline;

petiole 0.3–1.8 cm;

blade unlobed, narrowly ovate to elliptic, 0.3–2 × 0.3–1 cm, margins entire or inconspicuously denticulate, brownish oil cells lining veins abaxially.

in well-developed basal rosette and also cauline, 7–20 × 1.5–2.5 cm;

petiole 1–4(–8) cm;

blade pinnately lobed, terminal lobe oblanceolate to narrowly ovate, 2.5–5 × 1–2.5 cm, margins irregularly dentate, oil cells on abaxial surface inconspicuous.

Racemes

erect, elongating in fruit.

nodding, dense, mostly elongating after flowers open.

Flowers

opening at sunrise;

buds without free tips;

floral tube 0.4–0.5 mm, glabrous inside;

sepals 1–1.2 mm;

petals yellow, fading pale lavender, 1–1.5 mm;

stamens 4 (or 8), antisepalous, filaments 0.5 mm, anthers 0.5–0.7 mm, glabrous, when 8, then antipetalous ones smaller and abortive;

style 1.5 mm, stigma surrounded by anthers at anthesis.

opening at sunrise;

buds with conspicuous, subapical free tips 1–2 mm;

floral tube 3–5 mm, short-villous inside proximally;

sepals 9–12 mm;

petals bright yellow, with red dots at base, fading lavender, 12–18 mm;

stamens unequal, filaments of antisepalous stamens 6–8 mm, those of antipetalous ones 4–5 mm, anthers 4–6 mm, ciliate;

style 11–18 mm, stigma exserted beyond anthers at anthesis.

Capsules

spreading or reflexed, clavate, 4–10 mm;

pedicel 3–9 mm.

ascending or spreading, oblong-cylindrical, immature capsule to 35 mm;

pedicel 5–15 mm.

Seeds

0.8 mm.

not known.

2n

= 14.

Chylismia exilis

Chylismia confertiflora

Phenology Flowering Apr–Jun. Flowering Apr–May.
Habitat Calcareous sand, gypseous clay flats, juniper woodlands. Cinder soil.
Elevation 1000–1900 m. (3300–6200 ft.) 1300–1400 m. (4300–4600 ft.)
Distribution
from FNA
AZ; UT
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Chylismia exilis, known from Kane and San Juan counties in Utah and northern Coconino and Mohave counties in Arizona, is cryptic due to its small size. It may not be as rare as assumed, since it is difficult to spot in the field. P. H. Raven (1962, 1969) determined this species to be self-compatible and autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chylismia confertiflora is known only from the type locality on the east side and base of Vulcan’s Throne, Toroweap Valley, Grand Canyon National Monument in Mohave County. P. H. Raven (1962, 1969) assumed this species to be self-incompatible, based on the large flowers with the stigma elevated above the anthers.

A. Cronquist et al. (1997c) treated Chylismia confertiflora as part of C. brevipes, with a comment about the capsule dimensions, and indicated they consider the differences to be one end of the spectrum of a variation within C. brevipes. Although known from very few collections and sparse field data, P. H. Raven (1962, 1969) considered it to be distinct and to be most closely related to C. brevipes and to C. multijuga. He distinguished it from the latter by its larger flowers, nodding inflorescences, and large buds; and from the former by its glandular puberulent sepals, unequal stamens, and uniformly branched habit. In addition, the very restricted range of C. confertiflora is outside (to the east) of the range of C. brevipes. Chylismia multijuga grows within a few miles of the only know locality, but in this area C. confertiflora is very distinct from that species. Further collections and more detailed study of the overall morphological patterns as well as, perhaps, molecular data may clarify whether this is best considered to be an extremely restricted distinct species or a somewhat distinct outlier of the variable C. brevipes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
Synonyms Oenothera exilis, Camissonia exilis Oenothera confertiflorap., Camissonia confertiflora
Name authority (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007)
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