Chylismia exilis |
Chylismia |
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beeblossom, browneyes |
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Habit | Herbs annual, glandular puberulent and sparsely villous. | Herbs,usually annual, sometimes perennial, rarely biennial, usually caulescent. | ||||
Stems | slender, unbranched or branched, 10–20 cm. |
ascending to erect, usually branched. |
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Leaves | primarily cauline; petiole 0.3–1.8 cm; blade unlobed, narrowly ovate to elliptic, 0.3–2 × 0.3–1 cm, margins entire or inconspicuously denticulate, brownish oil cells lining veins abaxially. |
basal and cauline, cauline often reduced, basal often forming well-developed rosette, alternate; stipules absent; long-petiolate; blade often pinnately (rarely bipinnately) lobed, sometimes unlobed, or lateral lobes greatly reduced or absent, terminal lobe usually large, margins usually regularly or irregularly dentate to serrate, sometimes denticulate, serrulate, or entire, abaxial surface or margin with ± conspicuous, usually brown, oil cells. |
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Racemes | erect, elongating in fruit. |
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Inflorescences | racemes, erect or nodding. |
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Flowers | opening at sunrise; buds without free tips; floral tube 0.4–0.5 mm, glabrous inside; sepals 1–1.2 mm; petals yellow, fading pale lavender, 1–1.5 mm; stamens 4 (or 8), antisepalous, filaments 0.5 mm, anthers 0.5–0.7 mm, glabrous, when 8, then antipetalous ones smaller and abortive; style 1.5 mm, stigma surrounded by anthers at anthesis. |
bisexual, actinomorphic, buds usually erect, sometimes reflexed; floral tube deciduous (with sepals, petals, and stamens after anthesis), with basal nectary; sepals 4, reflexed singly; petals 4, usually yellow or white, often fading orange-red, sometimes lavender or purple, rarely cream, often with 1+ red dots near base; stamens usually 8, in 2 subequal series, rarely 4 in 1 series (usually in C. exilis), anthers versatile, pollen shed singly or in tetrads; ovary 4-locular, stigma usually entire and capitate, rarely conical-peltate and ± 4-lobed, surface unknown, probably wet and non-papillate. |
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Fruit | a capsule, straight or slightly curved, subterete and clavate or oblong-cylindrical, regularly loculicidal; pedicellate. |
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Capsules | spreading or reflexed, clavate, 4–10 mm; pedicel 3–9 mm. |
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Seeds | 0.8 mm. |
numerous, in 2 rows per locule, lenticular to narrowly ovoid to narrowly obovoid, finely pitted, with ± pronounced membranous margin when immature. |
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x |
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2n | = 14. |
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Chylismia exilis |
Chylismia |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Calcareous sand, gypseous clay flats, juniper woodlands. | |||||
Elevation | 1000–1900 m. (3300–6200 ft.) | |||||
Distribution |
AZ; UT |
w United States; nw Mexico |
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Discussion | Chylismia exilis, known from Kane and San Juan counties in Utah and northern Coconino and Mohave counties in Arizona, is cryptic due to its small size. It may not be as rare as assumed, since it is difficult to spot in the field. P. H. Raven (1962, 1969) determined this species to be self-compatible and autogamous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 16 (16 in the flora). Chylismia is distinguished from other genera formerly included in Camissonia by straight to arcuate (never twisted or curled) capsules on distinct pedicels and seeds in 2 rows per locule. R. A. Levin et al. (2004) included only one species each from Camissonia sects. Chylismia and Lignothera; the two formed a moderately supported branch, which led W. L. Wagner et al. (2007) to recognize Chylismia as a distinct genus. Chylismia is strongly supported in a sister relationship to the realigned Oenothera. This clade is in turn sister to Eulobus. Reproductive features include: self-incompatible (C. brevipes, C. claviformis, C. multijuga, C. munzii, and probably C. confertiflora, C. eastwoodiae, and C. parryi; P. H. Raven 1962, 1969) or self-compatible; flowers diurnal, outcrossing and pollinated by mostly oligolectic bees or autogamous, or opening one to two hours before sunset (in one subspecies of C. claviformis and the two species of sect. Lignothera); the evening-opening subspecies of C. claviformis pollinated mostly by oligolectic bees and moths, C. cardiophylla mainly by small moths, and C. arenaria, with its long floral tubes, by hawkmoths (E. G. Linsley et al. 1963, 1963b, 1964). Most species are diploid (2n = 14) but there are occasional tetraploids (2n = 28); floating translocations are relatively common (Raven 1962, 1969). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Oenothera exilis, Camissonia exilis | Oenothera, Camissonia section chylismia, Oenothera section chylismia, Oenothera subg. chylismia | ||||
Name authority | (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007) | (Torrey & A. Gray) Nuttall ex Raimann in H. G. A. Engler and K. Prantl: Nat. Pflanzenfam. 96[III,7]: 217. (1893) | ||||
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