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naked-stem beeblossom, Paiute suncup

Habit Herbs annual, succulent, glabrous, glandular puberulent, or villous proximally. Herbs annual, strigillose, villous, or glandular puberulent.
Stems

unbranched or branched from base, 3–30 cm.

usually unbranched, sometimes branched from base, 3–45 cm.

Leaves

primarily in basal rosette and also cauline;

petiole 0.5–8 cm;

blade usually not pinnately lobed or, if so, then lateral lobes greatly reduced, terminal lobe oblanceolate to cordate, 0.8–7.5 × 0.4–3 cm, margins entire or sparsely denticulate, pale brown oil cells lining veins abaxially.

primarily in basal rosette, cauline poorly developed or absent, 1–18 × 0.5–3.5 cm;

petiole 0.5–6.5 cm;

blade pinnately lobed or lateral lobes greatly reduced or absent, sometimes mixed on same plant, terminal lobe narrowly ovate to ovate or elliptic, 1–6.5 × 0.5–3.3 cm, margins irregularly dentate to subentire, oil cells on abaxial surface inconspicuous or conspicuous, pale yellowish brown or dark brown.

Racemes

nodding, elongating after anthesis.

nodding, elongating in fruit.

Flowers

opening at sunrise;

buds without subapical free tips;

floral tube 2–4.5 mm, villous inside proximally;

sepals 3–8 mm;

petals bright yellow, with red dots near base, fading pale yellow or yellowish orange, 5.5–9 mm;

stamens unequal, filaments of antisepalous stamens 3–8 mm, those of antipetalous ones 2.8–5.5 mm, anthers 2–4 mm, ciliate;

style 10–17 mm, stigma exserted beyond anthers at anthesis.

opening at sunrise;

buds with or without subapical free tips less than 1 mm;

floral tube 1–4 mm, sparsely villous or glabrous inside;

sepals 1.2–5 mm;

petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 1.5–5.5(–8) mm;

stamens unequal, filaments of antisepalous stamens 1.2–6 mm, those of antipetalous ones 0.5–4 mm, anthers 1–2.5 mm, ciliate or glabrous;

style 3–11 mm, stigma surrounded by anthers at anthesis.

Capsules

erect, spreading, or slightly reflexed, clavate, 18–40 mm;

pedicel 4–28 mm.

ascending, clavate, 8–50 mm;

pedicel 4–20 mm.

Seeds

1.2–1.7 mm.

1–2 mm.

2n

= 14.

Chylismia eastwoodiae

Chylismia scapoidea

Phenology Flowering Apr–Jun.
Habitat Clay flats, on gray, alkaline, marine-deposited gumbo, sandy draws.
Elevation 1200–1800 m. (3900–5900 ft.)
Distribution
from FNA
CO; UT
[BONAP county map]
w United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Chylismia eastwoodiae is known from Mesa County, Colorado, and from Emery County south to San Juan County, Utah. P. H. Raven (1962, 1969) suspected this species to be self-incompatible, but did not have data to make the determination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-compatible and primarily autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Oil cells lining veins of leaf blades abaxially usually pale yellowish brown, inconspicuous, rarely darker; leaf blades usually unlobed, rarely pinnately lobed with few, small lateral lobes, margins subentire.
→ 2
2. Capsules (10–)15–30 mm; petals 1.7–5 mm.
subsp. scapoidea
2. Capsules 25–50 mm; petals 1.5–2 mm.
subsp. macrocarpa
1. Oil cells lining veins of leaf blades abaxially dark brown, conspicuous; leaf blades pinnately lobed, lobes well developed or with few, small lateral lobes or unlobed, margins dentate.
→ 3
3. Petals 1.8–4 mm; lateral lobes of leaves often few or absent, sometimes well developed; capsules 8–20(–28) mm.
subsp. brachycarpa
3. Petals 4–5.5(–8) mm; lateral lobes of leaves usually well developed, sometimes poorly developed; capsules 16–38 mm.
subsp. utahensis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. specicola, C. walkeri
Subordinate taxa
C. scapoidea subsp. brachycarpa, C. scapoidea subsp. macrocarpa, C. scapoidea subsp. scapoidea, C. scapoidea subsp. utahensis
Synonyms Oenothera scapoidea var. eastwoodiae, Camissonia eastwoodiae, O. eastwoodiae Oenothera scapoidea, Camissonia scapoidea, O. brevipes var. scapoidea
Name authority (Munz) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007) (Torrey & A. Gray) Nuttall ex Raimann in H. G. A. Engler and K. Prantl: Nat. Pflanzenfam. 96[III,7]: 217. (1893)
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