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redclay suncup

Habit Herbs annual, densely villous and strigillose, glandular puberulent on distal parts. Herbs annual, sparsely to densely villous throughout or, sometimes, glabrate distally.
Stems

well branched, 15–50 cm.

often intricately branched, 5–80 cm.

Leaves

in well-developed basal rosette and also cauline, 7–20 × 1.5–2.5 cm;

petiole 1–4(–8) cm;

blade pinnately lobed, terminal lobe oblanceolate to narrowly ovate, 2.5–5 × 1–2.5 cm, margins irregularly dentate, oil cells on abaxial surface inconspicuous.

in poorly defined basal rosette and also cauline;

petiole 0.3–3.8 cm;

blade usually unlobed, very rarely pinnately lobed with few, small lateral lobes, ovate to elliptic, margins sparsely denticulate to subentire, pale or dark brown oil cells lining veins abaxially.

Racemes

nodding, dense, mostly elongating after flowers open.

nodding, with intricate, filiform branches, elongating in fruit.

Flowers

opening at sunrise;

buds with conspicuous, subapical free tips 1–2 mm;

floral tube 3–5 mm, short-villous inside proximally;

sepals 9–12 mm;

petals bright yellow, with red dots at base, fading lavender, 12–18 mm;

stamens unequal, filaments of antisepalous stamens 6–8 mm, those of antipetalous ones 4–5 mm, anthers 4–6 mm, ciliate;

style 11–18 mm, stigma exserted beyond anthers at anthesis.

opening at sunrise;

buds without free tips;

floral tube 0.5–2 mm, glabrous or villous inside;

sepals 1.5–4 mm;

petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 2–7 mm;

stamens unequal, filaments of antisepalous stamens 1.7–3.5 mm, those of antipetalous ones 1.2–2.5 mm, anthers 0.9–1.2 mm, glabrous;

style 4–9 mm, stigma exserted beyond anthers at anthesis.

Capsules

ascending or spreading, oblong-cylindrical, immature capsule to 35 mm;

pedicel 5–15 mm.

erect or ascending, clavate, 4–10 mm;

pedicel 4–20 mm.

Seeds

not known.

0.7–1.2 mm.

2n

= 14.

Chylismia confertiflora

Chylismia parryi

Phenology Flowering Apr–May. Flowering May–Jun(–Sep).
Habitat Cinder soil. Red clay and sand slopes weathered from red (freshwater-deposited) sandstone cliffs, with Juniperus or Larrea tridentata.
Elevation 1300–1400 m. (4300–4600 ft.) 800–1300 m. (2600–4300 ft.)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
AZ; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Chylismia confertiflora is known only from the type locality on the east side and base of Vulcan’s Throne, Toroweap Valley, Grand Canyon National Monument in Mohave County. P. H. Raven (1962, 1969) assumed this species to be self-incompatible, based on the large flowers with the stigma elevated above the anthers.

A. Cronquist et al. (1997c) treated Chylismia confertiflora as part of C. brevipes, with a comment about the capsule dimensions, and indicated they consider the differences to be one end of the spectrum of a variation within C. brevipes. Although known from very few collections and sparse field data, P. H. Raven (1962, 1969) considered it to be distinct and to be most closely related to C. brevipes and to C. multijuga. He distinguished it from the latter by its larger flowers, nodding inflorescences, and large buds; and from the former by its glandular puberulent sepals, unequal stamens, and uniformly branched habit. In addition, the very restricted range of C. confertiflora is outside (to the east) of the range of C. brevipes. Chylismia multijuga grows within a few miles of the only know locality, but in this area C. confertiflora is very distinct from that species. Further collections and more detailed study of the overall morphological patterns as well as, perhaps, molecular data may clarify whether this is best considered to be an extremely restricted distinct species or a somewhat distinct outlier of the variable C. brevipes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chylismia parryi is known from northwestern Arizona (Coconino to Mohave counties) and southwestern Utah (Beaver to Washington counties), and is apparently disjunct to San Juan County, Utah. It is outcrossing and, perhaps, self-incompatible (P. H. Raven 1962, 1969). There are two morphological forms of this species. Raven (1962) noted that a later flowering form has narrower, smaller leaves, and less overall pubescence. It is not clear what these represent, but Raven (1962) made the combination Oenothera parryi forma tenuissima (M. E. Jones) P. H. Raven for it. He later (Raven 1969) noted that these plants did not seem to merit formal recognition, without any discussion.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. scapoidea, C. specicola, C. walkeri
Synonyms Oenothera confertiflorap., Camissonia confertiflora Oenothera parryi, Camissonia parryi, C. tenuissima, O. scapoidea var. parryi, O. tenuissima
Name authority (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 207. (2007) (S. Watson) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma)
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