Chylismia confertiflora
Chylismia
beeblossom, browneyes
well branched, 15–50 cm.
ascending to erect, usually branched.
in well-developed basal rosette and also cauline, 7–20 × 1.5–2.5 cm;
petiole 1–4(–8) cm;
blade pinnately lobed, terminal lobe oblanceolate to narrowly ovate, 2.5–5 × 1–2.5 cm, margins irregularly dentate, oil cells on abaxial surface inconspicuous.
basal and cauline, cauline often reduced, basal often forming well-developed rosette, alternate;
stipules absent; long-petiolate;
blade often pinnately (rarely bipinnately) lobed, sometimes unlobed, or lateral lobes greatly reduced or absent, terminal lobe usually large, margins usually regularly or irregularly dentate to serrate, sometimes denticulate, serrulate, or entire, abaxial surface or margin with ± conspicuous, usually brown, oil cells.
nodding, dense, mostly elongating after flowers open.
racemes, erect or nodding.
opening at sunrise;
buds with conspicuous, subapical free tips 1–2 mm;
floral tube 3–5 mm, short-villous inside proximally;
sepals 9–12 mm;
petals bright yellow, with red dots at base, fading lavender, 12–18 mm;
stamens unequal, filaments of antisepalous stamens 6–8 mm, those of antipetalous ones 4–5 mm, anthers 4–6 mm, ciliate;
style 11–18 mm, stigma exserted beyond anthers at anthesis.
bisexual, actinomorphic, buds usually erect, sometimes reflexed;
floral tube deciduous (with sepals, petals, and stamens after anthesis), with basal nectary;
sepals 4, reflexed singly;
petals 4, usually yellow or white, often fading orange-red, sometimes lavender or purple, rarely cream, often with 1+ red dots near base;
stamens usually 8, in 2 subequal series, rarely 4 in 1 series (usually in C. exilis), anthers versatile, pollen shed singly or in tetrads;
ovary 4-locular, stigma usually entire and capitate, rarely conical-peltate and ± 4-lobed, surface unknown, probably wet and non-papillate.
a capsule, straight or slightly curved, subterete and clavate or oblong-cylindrical, regularly loculicidal;
pedicellate.
ascending or spreading, oblong-cylindrical, immature capsule to 35 mm;
pedicel 5–15 mm.
not known.
numerous, in 2 rows per locule, lenticular to narrowly ovoid to narrowly obovoid, finely pitted, with ± pronounced membranous margin when immature.
Chylismia confertiflora
Chylismia
Chylismia confertiflora is known only from the type locality on the east side and base of Vulcan’s Throne, Toroweap Valley, Grand Canyon National Monument in Mohave County. P. H. Raven (1962, 1969) assumed this species to be self-incompatible, based on the large flowers with the stigma elevated above the anthers.
A. Cronquist et al. (1997c) treated Chylismia confertiflora as part of C. brevipes, with a comment about the capsule dimensions, and indicated they consider the differences to be one end of the spectrum of a variation within C. brevipes. Although known from very few collections and sparse field data, P. H. Raven (1962, 1969) considered it to be distinct and to be most closely related to C. brevipes and to C. multijuga. He distinguished it from the latter by its larger flowers, nodding inflorescences, and large buds; and from the former by its glandular puberulent sepals, unequal stamens, and uniformly branched habit. In addition, the very restricted range of C. confertiflora is outside (to the east) of the range of C. brevipes. Chylismia multijuga grows within a few miles of the only know locality, but in this area C. confertiflora is very distinct from that species. Further collections and more detailed study of the overall morphological patterns as well as, perhaps, molecular data may clarify whether this is best considered to be an extremely restricted distinct species or a somewhat distinct outlier of the variable C. brevipes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 16 (16 in the flora).
Chylismia is distinguished from other genera formerly included in Camissonia by straight to arcuate (never twisted or curled) capsules on distinct pedicels and seeds in 2 rows per locule. R. A. Levin et al. (2004) included only one species each from Camissonia sects. Chylismia and Lignothera; the two formed a moderately supported branch, which led W. L. Wagner et al. (2007) to recognize Chylismia as a distinct genus. Chylismia is strongly supported in a sister relationship to the realigned Oenothera. This clade is in turn sister to Eulobus. Reproductive features include: self-incompatible (C. brevipes, C. claviformis, C. multijuga, C. munzii, and probably C. confertiflora, C. eastwoodiae, and C. parryi; P. H. Raven 1962, 1969) or self-compatible; flowers diurnal, outcrossing and pollinated by mostly oligolectic bees or autogamous, or opening one to two hours before sunset (in one subspecies of C. claviformis and the two species of sect. Lignothera); the evening-opening subspecies of C. claviformis pollinated mostly by oligolectic bees and moths, C. cardiophylla mainly by small moths, and C. arenaria, with its long floral tubes, by hawkmoths (E. G. Linsley et al. 1963, 1963b, 1964). Most species are diploid (2n = 14) but there are occasional tetraploids (2n = 28); floating translocations are relatively common (Raven 1962, 1969).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Floral tubes 0.4–9 mm; pollen shed singly; leaves basal and cauline, usually with well-developed basal rosettes, blades usually pinnately or bipinnately lobed, lateral lobes sometimes greatly reduced or absent; plants usually annual, sometimes perennial, rarely biennial. | sect. Chylismia |
1. Floral tubes 4.5–40 mm; pollen shed in tetrads; leaves cauline, blades unlobed; plants usually perennial, sometimes annual. | sect. Lignothera |
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