Chylismia claviformis |
Chylismia scapoidea |
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brown-eyed primrose, browneyes, clavate fruit primrose |
naked-stem beeblossom, Paiute suncup |
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Habit | Herbs annual, glabrous, strigillose, glandular puberulent, or, sometimes, villous. | Herbs annual, strigillose, villous, or glandular puberulent. | ||||||||||||||||||||||||||||||||||||||||||||||||
Stems | branched mostly from base, 3–70 cm. |
usually unbranched, sometimes branched from base, 3–45 cm. |
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Leaves | primarily in basal rosette, cauline reduced or absent, 1.5–20 × 0.3–3.5 cm; petiole 0.7–12 cm; blade usually pinnately lobed, sometimes lateral lobes poorly developed or absent, terminal lobe usually narrowly ovate to lanceolate, sometimes cordate or subcordate, 0.8–9 × 0.2–4.5 cm, margins dentate, sinuate-dentate, or serrate, brown oil cells conspicuously lining veins abaxially. |
primarily in basal rosette, cauline poorly developed or absent, 1–18 × 0.5–3.5 cm; petiole 0.5–6.5 cm; blade pinnately lobed or lateral lobes greatly reduced or absent, sometimes mixed on same plant, terminal lobe narrowly ovate to ovate or elliptic, 1–6.5 × 0.5–3.3 cm, margins irregularly dentate to subentire, oil cells on abaxial surface inconspicuous or conspicuous, pale yellowish brown or dark brown. |
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Racemes | nodding, elongating after anthesis. |
nodding, elongating in fruit. |
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Flowers | opening at sunset or sunrise; buds with or without subapical or apical free tips; floral tube 2–6.5 mm, villous inside proximally; sepals 2–8 mm; petals pale to bright yellow or white, sometimes red- or purple-dotted near base, fading purple, sometimes red or orange, or not changing color, 1.5–8 mm; stamens subequal, filaments 1.5–5.5 mm, anthers 1.5–6 mm, ciliate; style 5–16 mm, stigma exserted beyond anthers at anthesis. |
opening at sunrise; buds with or without subapical free tips less than 1 mm; floral tube 1–4 mm, sparsely villous or glabrous inside; sepals 1.2–5 mm; petals bright yellow, often with red dots near base, fading pale yellow or yellowish orange, 1.5–5.5(–8) mm; stamens unequal, filaments of antisepalous stamens 1.2–6 mm, those of antipetalous ones 0.5–4 mm, anthers 1–2.5 mm, ciliate or glabrous; style 3–11 mm, stigma surrounded by anthers at anthesis. |
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Capsules | ascending to spreading, clavate, 8–40 mm; pedicel 4–40 mm. |
ascending, clavate, 8–50 mm; pedicel 4–20 mm. |
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Seeds | 0.6–1.5 mm. |
1–2 mm. |
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Chylismia claviformis |
Chylismia scapoidea |
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Distribution | w United States; nw Mexico
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w United States
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Discussion | Subspecies 11 (10 in the flora). P. H. Raven (1962) subdivided this species into 12 subspecies and, subsequently (1969), he combined two of them. The latter approach is used here. Only subsp. wigginsii P. H. Raven does not occur in the United States; its narrow range is restricted to northern Baja California. Raven (1962, 1969) determined this species to be self-incompatible. Chylismia claviformis is the most complex and, along with C. scapoidea, the most widely distributed species of the genus. The central part of its geographical range is occupied by five closely related white-petaled subspecies (aurantiaca, claviformis, funerea, integrior, and peeblesii) that are very similar morphologically. South of this area four additional subspecies occur, all yellow-petaled (peirsonii, rubescens, wigginsii, and yumae). These four subspecies have sepals and petal color similar to those of C. brevipes, and P. H. Raven (1962, 1969) thought it likely that they were derived following hybridization between that species and one of the white-petaled populations of C. claviformis. North of the range of the white-petaled subspecies are found two additional yellow-petaled subspecies (cruciformis and lancifolia). Most populations of subsp. cruciformis consist of plants in which the flowers open in the early morning; in all other subspecies the flowers open in the late afternoon (Raven 1962, 1969). The following key will separate them, but there are many intergrades among the subspecies so that not all specimens will be easily identified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 4 (4 in the flora). P. H. Raven (1962, 1969) determined this species to be self-compatible and primarily autogamous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Oenothera claviformis, Camissonia claviformis, C. scapoidea var. claviformis, O. scapoidea var. claviformis | Oenothera scapoidea, Camissonia scapoidea, O. brevipes var. scapoidea | ||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Torrey & Frémont) A. Heller: Muhlenbergia 2: 105. (1906) — (as Chylisma clavaeformis) | (Torrey & A. Gray) Nuttall ex Raimann in H. G. A. Engler and K. Prantl: Nat. Pflanzenfam. 96[III,7]: 217. (1893) | ||||||||||||||||||||||||||||||||||||||||||||||||
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