Chylismia claviformis subsp. integrior |
Onagraceae subfam. onagroideae |
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browneyes, clavate fruit primrose |
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Habit | Herbs strigillose proximally, usually also glandular puberulent distally, or glabrate. | |
Stems | 6–70 cm. |
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Leaves | blade sometimes purple-dotted, lateral lobes often absent, sometimes present but poorly developed, terminal lobe narrowly ovate to ovate or subcordate, to 7 × 3 cm, margins serrate. |
stipules present or absent. |
Flowers | opening at sunset; buds usually without free tips, rarely with apical free tips less than 1 mm; floral tube orange-brown inside, 3–6 mm; petals white, sometimes purple-dotted near base, fading purple, 4.5–8 mm. |
floral tube present or, rarely, absent; sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens; petals yellow, white, pink, red, rarely in combination. |
x |
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2n | = 14. |
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Chylismia claviformis subsp. integrior |
Onagraceae subfam. onagroideae |
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Phenology | Flowering Apr–Jun. | |
Habitat | Dry flats, with Artemisia tridentata, Ericameria, or Juniperus, on banks and flats. | |
Elevation | 1100–2200 m. (3600–7200 ft.) | |
Distribution |
CA; NV; OR |
North America; Mexico; Central America; South America; West Indies; Eurasia; Pacific Islands (New Zealand, Society Islands); Australia |
Discussion | Subspecies integrior is known from southern Harney County, Oregon, southward nearly throughout Nevada to Esmeralda and Lincoln counties and to Mono and northeastern Inyo counties, California, and often forms large colonies on banks and flats in Great Basin Desert. It intergrades with subspp. aurantiaca and cruciformis, and hybridizes with Chylismia brevipes subsp. brevipes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 21, species 582 (16 genera, 246 species in the flora). Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
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Synonyms | Oenothera claviformis subsp. integrior, Camissonia claviformis subsp. integrior, C. claviformis var. purpurascens, O. claviformis var. purpurascens, O. scapoidea var. purpurascens | |
Name authority | (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 206. (2007) | W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 41. (2007) |
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