Chylismia claviformis subsp. cruciformis |
Onagraceae subfam. onagroideae |
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browneyes, cross-flower evening primrose, cruciform evening-primrose |
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Habit | Herbs strigillose or glandular puberulent proximally, glandular puberulent or glabrous distally. | |
Stems | 3–55 cm. |
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Leaves | blade lateral lobes well developed, few to numerous, terminal lobe narrowly ovate to subcordate, to 8 × to 4 cm, margins serrate-dentate. |
stipules present or absent. |
Flowers | opening at sunrise; buds without free tips, sometimes with apical free tips less than 1 mm; floral tube yellow or orange-brown inside, 2–6.5 mm; petals bright yellow, sometimes red-dotted in proximal 1/2, often fading purple, 2.5–8 mm. |
floral tube present or, rarely, absent; sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens; petals yellow, white, pink, red, rarely in combination. |
x |
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2n | = 14. |
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Chylismia claviformis subsp. cruciformis |
Onagraceae subfam. onagroideae |
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Phenology | Flowering Mar–May. | |
Habitat | Sandy or clay flats and slopes, with Artemisia tridentata, Ericameria, Grayia spinosa, or Purshia tridentata. | |
Elevation | 600–1500 m. (2000–4900 ft.) | |
Distribution |
CA; ID; NV; OR |
North America; Mexico; Central America; South America; West Indies; Eurasia; Pacific Islands (New Zealand, Society Islands); Australia |
Discussion | Subspecies cruciformis is known from Lassen County, California, western Canyon and Owyhee counties, Idaho, Harney, Lake, and Malheur counties, Oregon, and central and southern Washoe County, Nevada. It intergrades with subsp. integrior. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 21, species 582 (16 genera, 246 species in the flora). Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
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Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Oenothera cruciformis, Camissonia claviformis subsp. cruciformis, C. claviformis var. cruciformis, Chylisma cruciformis, C. scapoidea var. cruciformis, O. claviformis subsp. citrina, O. claviformis subsp. cruciformis, O. claviformis var. cruciformis | |
Name authority | (Kellogg) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 206. (2007) | W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 41. (2007) |
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