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heart leaf primrose, heartleaf suncup

Habit Herbs annual or perennial, villous and glandular puberulent. Herbs perennial, base sometimes woody, glabrous or sparsely villous proximally.
Stems

usually well branched, forming bushy habit, 20–100 cm.

with several divergent branches from base, 10–50 cm.

Leaves

cauline, mostly toward base;

petiole (0.7–)2.5–7.5 cm;

blade cordate-ovate to -orbiculate, 2.5–7.5 × 2.3–5.5 cm, smaller distally, margins erose-dentate.

primarily in basal rosette and cauline, 3–20 × 0.7–2.5 cm;

petiole 0.5–4 cm;

blade pinnately or bipinnately lobed, terminal lobe ovate to elliptic, 0.4–2.5 × 0.3–1.5 cm, margins irregularly serrate, dark brown oil cells prominently lining veins abaxially.

Racemes

nodding, congested.

erect, elongating after anthesis.

Flowers

floral tube 4.5–14 mm, villous inside;

sepals 3–9 mm;

petals yellow, 3–12 mm;

filaments 1–3 mm, anthers 2–4 mm;

style 8–23 mm, stigma surrounded by or exserted just beyond anthers at anthesis.

opening at sunrise;

buds individually reflexed, with apical free tips less than 1 mm;

floral tube 1.5–2 mm, glabrous inside;

sepals 2–5 mm;

petals bright yellow, with red dots near base, fading pale lavender, 2–6 mm;

stamens unequal, filaments of antisepalous stamens 1.5–3 mm, those of antipetalous ones 1–2 mm, anthers 1.2–2 mm, glabrous;

style 4–7 mm, stigma surrounded by anthers at anthesis.

Capsules

ascending, cylindrical, 20–55 mm;

pedicel 1–18 mm.

spreading to ascending, oblong-cylindrical, 8–20 mm;

pedicel 6–10 mm.

Seeds

0.5–0.7 mm.

0.6–1 mm.

Chylismia cardiophylla

Chylismia specicola

Distribution
sw United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Arizona
[BONAP county map]
Discussion

Subspecies 3 (2 in the flora).

P. H. Raven (1962, 1969) determined this species to be self-compatible, but primarily outcrossing. Subspecies cedrosensis (Greene) W. L. Wagner & Hoch, occurs in Baja California and adjacent Sonora, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

P. H. Raven (1962, 1969) thought that this species is most likely self-compatible but primarily outcrossing.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs villous, sometimes also glandular puberulent; floral tubes 4.5–12 mm.
subsp. cardiophylla
1. Herbs glandular puberulent throughout, often also sparsely villous; floral tubes 9–14 mm.
subsp. robusta
1. Leaf blades and styles glabrous; herbs ± woody at base.
subsp. specicola
1. Leaf blades and styles villous, at least at base; herbs not woody.
subsp. hesperia
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Lignothera Onagraceae > subfam. Onagroideae > tribe Onagreae > Chylismia > sect. Chylismia
Sibling taxa
C. arenaria, C. atwoodii, C. brevipes, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. specicola, C. walkeri
C. arenaria, C. atwoodii, C. brevipes, C. cardiophylla, C. claviformis, C. confertiflora, C. eastwoodiae, C. exilis, C. heterochroma, C. megalantha, C. multijuga, C. munzii, C. parryi, C. scapoidea, C. walkeri
Subordinate taxa
C. cardiophylla subsp. cardiophylla, C. cardiophylla subsp. robusta
C. specicola subsp. hesperia, C. specicola subsp. specicola
Synonyms Oenothera cardiophylla, Camissonia cardiophylla Oenothera specicola, Camissonia specicola
Name authority (Torrey) Small: Bull. Torrey Bot. Club 23: 193. (1896) — (as Chylisma) (P. H. Raven) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 208. (2007)
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