Chylismia brevipes subsp. pallidula |
Onagraceae tribe Onagreae |
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pale sun cup |
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Habit | Herbs slender, usually branched distally, usually strigillose, sometimes also villous proximally. | Herbs (annual or perennial), [shrubs]. |
Leaves | alternate or basal; stipules absent. |
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Flowers | buds not individually reflexed, without free tips or with apical free tips less than 1 mm; floral tube yellow inside, 4–5 mm; petals often with red dots near base, fading yellow to orange, 7–12 mm; anthers 4–6 mm. |
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous; stamens 2 times as many, or rarely as many, as sepals; pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera). |
Fruit | a dry capsule, usually dehiscent, sometimes indehiscent. |
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Capsules | 20–42 mm; pedicel 2–10 mm. |
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Seeds | few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella). |
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2n | = 14. |
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Chylismia brevipes subsp. pallidula |
Onagraceae tribe Onagreae |
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Phenology | Flowering Mar–May. | |
Habitat | Dry flats, desert pavement, with Ambrosia dumosa and Larrea. | |
Elevation | 70–1100 m. (200–3600 ft.) | |
Distribution |
AZ; CA; NV; UT |
North America; Mexico; Central America; South America; West Indies |
Discussion | Subspecies pallidula occurs in the Sonoran and Mojave deserts from Washington County, Utah, to southeastern Inyo County, California, south to Riverside County, California, Mohave County, Arizona, and Clark County, Nevada. It hybridizes with Chylismia claviformis subspp. aurantiaca and peeblesii, as well as with C. multijuga. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 13, species 265 (12 genera, 199 species in the flora). Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Oenothera brevipes var. pallidula, Camissonia brevipes subsp. pallidula, O. brevipes subsp. pallidula, O. pallidula | |
Name authority | (Munz) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 206. (2007) | Dumortier: Fl. Belg., 89. (1827) |
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