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riverside spineflower, Xantus' spineflower

five-tooth spineflower, Watson's spineflower

Habit Plants erect to infrequently spreading, (0.3–)0.5–2.5(–3) × 0.5–3(–5) dm, thinly pubescent. Plants spreading to erect, 0.2–1(–1.5) × 0.2–1(–1.5) dm, densely canescent-strigose.
Leaves

basal or nearly so;

petiole 1–2(–3) cm;

blade oblong or oblong-ovate to ovate, 0.3–1(–1.5) × 0.3–0.8(–1) cm, thinly pubescent adaxially, densely tomentose abaxially.

basal or nearly so;

petiole (0.5–)1–2.5(–3) cm;

blade oblanceolate, (0.3–)0.5–1.5(–2) × 0.2–0.4(–0.5) cm, thinly floccose to sparsely tomentose.

Inflorescences

mostly flat-topped and openly branched, usually reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, oblong-ovate to ovate, 0.3–0.8 cm × 2–6 mm, becoming sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.4 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

with involucres in small clusters 0.5–1 cm diam., greenish to reddish;

bracts 2, with laminar ones oblanceolate, (0.5–)0.8–1.5(–2) cm × (1–)2–4 mm, those at distal nodes becoming sessile, reduced and scalelike, linear-lanceolate, acicular, awns slightly curved, 0.5–1 mm.

Involucres

in open clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4.5 mm, not corrugate, without scarious or membranous margins, thinly to densely pubescent;

teeth spreading, unequal, 0.7–1.5 mm, 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.5–1 mm.

1, green, cylindric, 5-ribbed, 3–4.5 mm, finely corrugate, pubescent;

teeth 5, erect, unequal, with leaflike, narrowly lanceolate, 2–6 mm anterior tooth, others linear, 1–2 mm;

awn uncinate, 0.4–0.8(–1) mm.

Flowers

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 2/3 their length, monomorphic to slightly dimorphic, narrowly oblanceolate, rounded apically, those of outer whorl occasionally slightly broader and longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

1, included to slightly exserted;

perianth yellow, cylindric, 1.5–2.5 mm, thinly pubescent abaxially;

tepals connate ca. 2/3 their length, monomorphic, oblong, acute, entire apically, mostly erect;

stamens 3 or 9, slightly exserted;

filaments distinct, 0.8–1 mm, glabrous;

anthers yellow, ovate, 0.2–0.3 mm.

Achenes

brown, lenticular, 4–4.5 mm.

brown, lenticular, 2.5–3 mm.

Chorizanthe xanti

Chorizanthe watsonii

Phenology Flowering Apr–Aug.
Habitat Sandy to gravelly flats and slopes, mixed grassland, saltbush and sagebrush communities, pinyon-juniper woodlands
Elevation 300-2400 m (1000-7900 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; ID; NV; OR; UT; WA
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[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe watsonii is widely distributed in the cold desert of the Great Basin and in the northern part of the warmer Mojave Desert. Plants in the northern part of the range (especially on the Palouse Prairie of south-eastern Washington) usually have three stamens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres thinly pubescent
var. xanti
1. Involucres densely white-pubescent
var. leucotheca
Source FNA vol. 5, p. 463. FNA vol. 5, p. 468.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. wheeleri, C. xanti
Subordinate taxa
C. xanti var. leucotheca, C. xanti var. xanti
Name authority S. Watson: Proc. Amer. Acad. Arts 12: 272. (1877) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 199. (1870)
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