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potbellied spineflower, Priest Valley spineflower

Santa Barbara spineflower, Wheeler's spineflower

Habit Plants spreading and diffuse, (0.5–)1–5 × 1–5(–7) dm, pubescent. Plants erect to spreading, 0.5–2(–2.5) × 1–2 dm, thinly pubescent.
Leaves

basal;

petiole 0.3–1(–1.5) cm;

blade oblanceolate, (0.5–)1–3(–4) × (0.2–)0.4–1(–1.2) cm, thinly pubescent.

basal;

petiole 0.5–3 cm;

blade elliptic to oblong, 0.5–2 × 0.2–0.6 cm, thinly pubescent adaxially, tomentose abaxially.

Inflorescences

with involucres in open clusters 2–6 cm diam., greenish or reddish;

bracts 2–3 at proximal node, usually leaflike, often with whorl of sessile bracts about midstem, oblanceolate to elliptic, 0.5–1.5 cm × 1–4 mm, gradually becoming reduced, linear-lanceolate, 0.4–1.2(–1.5) cm × 1.5–5 mm, at distal nodes scalelike, linear and aciculate, acerose, awns straight, 1–3 mm.

mostly flat-topped, openly branched, greenish to reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to leaf blades, short-petiolate, oblong, 0.5–1.2 cm × 2–4 mm, sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

Involucres

3–10+, greenish or reddish, urceolate, strongly ventricose basally, 4–4.5 mm, without scarious or membranous margins, corrugate, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight, mostly 2 mm, others uncinate, 0.5–1 mm.

in dense terminal clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 2–2.5 mm, corrugate, without scarious or membranous margins, thinly pubescent with stoutish, recurved hairs;

teeth spreading, unequal, 0.3–0.8(–1) mm, with 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.3–0.5 mm.

Flowers

exserted;

perianth bicolored with floral tube white to greenish yellow and tepals red to maroon, cylindric, 4–4.5 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, oblong, those of outer whorl spreading and recurved, slightly longer than those of inner whorl, broadly obcordate, slightly erose or at least wavy and rounded apically, those of inner whorl erect, narrower, fimbriate and truncate or somewhat 2-lobed apically, erect;

stamens 9, exserted;

filaments distinct, 3.5–4 mm, glabrous;

anthers pink to red or maroon, oblong, 1–1.3 mm.

exserted;

perianth white or rose or red with white lobes, cylindric, 2.5–3 mm, glabrous except for few scattered hairs ca. midlength along midrib abaxially;

tepals connate 1/2 their length, monomorphic to slightly dimorphic, oblong, rounded apically, those of outer whorl usually slightly broader and longer than those of inner whorl;

stamens 6, included;

filaments distinct, 1.5–2 mm, glabrous;

anthers pink to red, oblong, 0.3–0.4 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

brown, lenticular, 2.5–3 mm.

2n

= 40, 42, 44.

Chorizanthe ventricosa

Chorizanthe wheeleri

Phenology Flowering May–Sep. Flowering Apr–Jun.
Habitat Serpentine outcrops, mixed grassland communities, oak-pine woodlands Gravelly to rocky slopes, coastal scrub communities
Elevation 500-1000 m (1600-3300 ft) 0-400(-600) m (0-1300(-2000) ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe ventricosa is restricted to isolated outcrops of serpentine in the coastal mountain ranges of southeastern Monterey County and southern San Benito County south in western Fresno County to the Parkfield Grade area and in Cottonwood Pass of San Luis Obispo County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe wheeleri is a rare insular endemic known only from Santa Cruz and Santa Rosa islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 458. FNA vol. 5, p. 464.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. xanti
Synonyms C. palmeri var. ventricosa C. insularis
Name authority Goodman: Leafl. W. Bot. 2: 193, figs. 1, 2. (1939) S. Watson: Proc. Amer. Acad. Arts 12: 272. (1877)
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