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potbellied spineflower, Priest Valley spineflower

Habit Plants spreading and diffuse, (0.5–)1–5 × 1–5(–7) dm, pubescent. Plants prostrate to spreading or erect, mostly thinly pubescent.
Stems

sometimes disarticulating at each node.

Leaves

basal;

petiole 0.3–1(–1.5) cm;

blade oblanceolate, (0.5–)1–3(–4) × (0.2–)0.4–1(–1.2) cm, thinly pubescent.

blades linear to lanceolate, obovate, round, or spatulate.

Inflorescences

with involucres in open clusters 2–6 cm diam., greenish or reddish;

bracts 2–3 at proximal node, usually leaflike, often with whorl of sessile bracts about midstem, oblanceolate to elliptic, 0.5–1.5 cm × 1–4 mm, gradually becoming reduced, linear-lanceolate, 0.4–1.2(–1.5) cm × 1.5–5 mm, at distal nodes scalelike, linear and aciculate, acerose, awns straight, 1–3 mm.

bracts mostly 2, opposite, scalelike or if leaflike then similar to basal leaves only reduced, occasionally deciduous in early anthesis, with or without awns.

Involucres

3–10+, greenish or reddish, urceolate, strongly ventricose basally, 4–4.5 mm, without scarious or membranous margins, corrugate, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight, mostly 2 mm, others uncinate, 0.5–1 mm.

cylindric to narrowly turbinate, campanulate, or urceolate, occasionally ventricose basally, 3-,5-, or 6-toothed, with or without membranous or scarious margins;

teeth erect to spreading or divergent, connate at least 1/2 their length, typically shallow, mostly unequal, with alternating long and short awns, often with anterior one longest.

Flowers

exserted;

perianth bicolored with floral tube white to greenish yellow and tepals red to maroon, cylindric, 4–4.5 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, oblong, those of outer whorl spreading and recurved, slightly longer than those of inner whorl, broadly obcordate, slightly erose or at least wavy and rounded apically, those of inner whorl erect, narrower, fimbriate and truncate or somewhat 2-lobed apically, erect;

stamens 9, exserted;

filaments distinct, 3.5–4 mm, glabrous;

anthers pink to red or maroon, oblong, 1–1.3 mm.

1(–2), white to pink or rose, maroon or purple, or yellow, thinly pubescent at least along midribs abaxially;

stamens 3–9;

filaments adnate at base of floral tube or faucially;

filaments sometimes connate into short tube.

Achenes

brown, globose-lenticular, 3–3.5 mm.

brown, lenticular or globose-lenticular, or 3-gonous.

Seeds

embryo straight or rarely curved.

2n

= 40, 42, 44.

Chorizanthe ventricosa

Chorizanthe subg. Amphietes

Phenology Flowering May–Sep.
Habitat Serpentine outcrops, mixed grassland communities, oak-pine woodlands
Elevation 500-1000 m (1600-3300 ft)
Distribution
from FNA
CA
[BONAP county map]
w United States; nw Mexico; sw South America
Discussion

Chorizanthe ventricosa is restricted to isolated outcrops of serpentine in the coastal mountain ranges of southeastern Monterey County and southern San Benito County south in western Fresno County to the Parkfield Grade area and in Cottonwood Pass of San Luis Obispo County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 39 (31 in the flora).

Most species of subg. Amphietes are found in California. Of the others, one is known only from southernmost Peru to central Chile (Chorizanthe commissuralis J. Rémy), while the rest are known only from Baja California, Mexico. Those include C. inequalis S. Stokes, C. turbinata Wiggins, C. mutabilis Brandegee, C. rosulenta Reveal, C. pulchella Brandegee, C. flava Brandegee, and C. interposita Goodman. The latter is the only member of sect. Clastoscapa, the only section of subg. Amphietes not found in our flora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 458. FNA vol. 5, p. 450.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
Synonyms C. palmeri var. ventricosa
Name authority Goodman: Leafl. W. Bot. 2: 193, figs. 1, 2. (1939) Reveal & Hardham: Phytologia 66: 113. (1989)
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