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potbellied spineflower, Priest Valley spineflower

Monterey spineflower

Habit Plants spreading and diffuse, (0.5–)1–5 × 1–5(–7) dm, pubescent. Plants prostrate to ascending or erect, 0.5–2(–2.5) × 0.5–10 dm, grayish-villous.
Leaves

basal;

petiole 0.3–1(–1.5) cm;

blade oblanceolate, (0.5–)1–3(–4) × (0.2–)0.4–1(–1.2) cm, thinly pubescent.

basal;

petiole (0.5–)1–3(–4) cm;

blade oblanceolate, (0.5–)1–5(–7) × (0.3–)0.4–0.7(–1) cm, villous.

Inflorescences

with involucres in open clusters 2–6 cm diam., greenish or reddish;

bracts 2–3 at proximal node, usually leaflike, often with whorl of sessile bracts about midstem, oblanceolate to elliptic, 0.5–1.5 cm × 1–4 mm, gradually becoming reduced, linear-lanceolate, 0.4–1.2(–1.5) cm × 1.5–5 mm, at distal nodes scalelike, linear and aciculate, acerose, awns straight, 1–3 mm.

rather dense with secondary branches suppressed, grayish;

bracts 2, similar to leaf blades at proximal nodes only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–7 cm × 2–7 mm, awns 0.5–1.2 mm.

Involucres

3–10+, greenish or reddish, urceolate, strongly ventricose basally, 4–4.5 mm, without scarious or membranous margins, corrugate, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight, mostly 2 mm, others uncinate, 0.5–1 mm.

1, grayish, cylindric, often ventricose basally, 2–2.5(–3) mm, with distinct, white to pink or purple, scarious margins extending nearly full length of awn, corrugate, villous abaxially;

teeth spreading, equal, 0.5–1.5 mm;

awns uncinate with longer ones 2–3 mm and alternating with shorter (1–1.5 mm) ones.

Flowers

exserted;

perianth bicolored with floral tube white to greenish yellow and tepals red to maroon, cylindric, 4–4.5 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, oblong, those of outer whorl spreading and recurved, slightly longer than those of inner whorl, broadly obcordate, slightly erose or at least wavy and rounded apically, those of inner whorl erect, narrower, fimbriate and truncate or somewhat 2-lobed apically, erect;

stamens 9, exserted;

filaments distinct, 3.5–4 mm, glabrous;

anthers pink to red or maroon, oblong, 1–1.3 mm.

exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3.5 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, obovate to oblong, acute to truncate and erose apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, ovate, 0.3–0.4 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

dark brown, globose-lenticular, 2–2.5 mm.

2n

= 40, 42, 44.

Chorizanthe ventricosa

Chorizanthe pungens

Phenology Flowering May–Sep.
Habitat Serpentine outcrops, mixed grassland communities, oak-pine woodlands
Elevation 500-1000 m (1600-3300 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe ventricosa is restricted to isolated outcrops of serpentine in the coastal mountain ranges of southeastern Monterey County and southern San Benito County south in western Fresno County to the Parkfield Grade area and in Cottonwood Pass of San Luis Obispo County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white (rarely pinkish), scarious; plants prostrate to slightly ascending; coastal areas and adjacent inland valleys
var. pungens
1. Margins of involucres dark pinkish to purple, scarious; plants slightly ascending to erect; coastal mountains
var. hartwegiana
Source FNA vol. 5, p. 458. FNA vol. 5, p. 453.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. pungens var. hartwegiana, C. pungens var. pungens
Synonyms C. palmeri var. ventricosa
Name authority Goodman: Leafl. W. Bot. 2: 193, figs. 1, 2. (1939) Bentham: Trans. Linn. Soc. London 17: 419, plate 19, fig. 2. (1836)
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