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potbellied spineflower, Priest Valley spineflower

pink spineflower

Habit Plants spreading and diffuse, (0.5–)1–5 × 1–5(–7) dm, pubescent. Plants 1–6(–10) × 0.5–3(–5) dm, woolly-floccose.
Leaves

basal;

petiole 0.3–1(–1.5) cm;

blade oblanceolate, (0.5–)1–3(–4) × (0.2–)0.4–1(–1.2) cm, thinly pubescent.

basal and cauline;

petiole 0.1–0.5(–0.8) cm;

blade linear to narrowly oblanceolate, (1–)1.5–5 × 0.1–0.3 cm, thinly to densely floccose adaxially, densely tomentose abaxially.

Inflorescences

with involucres in open clusters 2–6 cm diam., greenish or reddish;

bracts 2–3 at proximal node, usually leaflike, often with whorl of sessile bracts about midstem, oblanceolate to elliptic, 0.5–1.5 cm × 1–4 mm, gradually becoming reduced, linear-lanceolate, 0.4–1.2(–1.5) cm × 1.5–5 mm, at distal nodes scalelike, linear and aciculate, acerose, awns straight, 1–3 mm.

strict, white to greenish, open;

bracts usually 2, opposite, rarely in whorls of 3–5, short-petiolate, acerose, similar to proximal leaf blades only reduced, 0.3–3 cm × 1–3 mm, awns straight, 0.5–1 mm.

Involucres

3–10+, greenish or reddish, urceolate, strongly ventricose basally, 4–4.5 mm, without scarious or membranous margins, corrugate, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight, mostly 2 mm, others uncinate, 0.5–1 mm.

usually congested in small terminal clusters of 1–3 at node of dichotomies, urceolate, ventricose basally, 3-angled, 6-ribbed, 3–4 mm, not corrugate, with conspicuous, white margins extending across sinuses, tomentose to floccose or glabrate with age, greenish to brownish;

teeth 6;

awns uncinate, 0.7–1.5 mm.

Flowers

exserted;

perianth bicolored with floral tube white to greenish yellow and tepals red to maroon, cylindric, 4–4.5 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, oblong, those of outer whorl spreading and recurved, slightly longer than those of inner whorl, broadly obcordate, slightly erose or at least wavy and rounded apically, those of inner whorl erect, narrower, fimbriate and truncate or somewhat 2-lobed apically, erect;

stamens 9, exserted;

filaments distinct, 3.5–4 mm, glabrous;

anthers pink to red or maroon, oblong, 1–1.3 mm.

1(–2), slightly exserted;

perianth white to rose, subcylindric, (1.5–)2.5–3 mm, densely pubescent abaxially;

tepals connate 2/3 their length, slightly dimorphic, entire and rounded apically, those of outer whorl obovate, those of inner whorl spatulate;

stamens slightly exserted;

filaments 1.5–2.5 mm, glabrous;

anthers pink to red, oval, 0.2–0.3 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

2.5–3 mm.

2n

= 40, 42, 44.

= 38, 40, (42), 80, 82, 84.

Chorizanthe ventricosa

Chorizanthe membranacea

Phenology Flowering May–Sep. Flowering Apr–Jul.
Habitat Serpentine outcrops, mixed grassland communities, oak-pine woodlands Sandy to gravelly or rocky flats and slopes, mixed grassland and chaparral communities, oak-pine woodlands
Elevation 500-1000 m (1600-3300 ft) 40-1400(-1600) m (100-4600(-5200) ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe ventricosa is restricted to isolated outcrops of serpentine in the coastal mountain ranges of southeastern Monterey County and southern San Benito County south in western Fresno County to the Parkfield Grade area and in Cottonwood Pass of San Luis Obispo County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe membranacea has long been considered an isolated element among the spineflowers. The strict, upright habit, numerous basal and cauline leaves, and broad, continuous, membranous margins of the involucre all reflect that isolation. Pink spineflower is widespread and often locally common in the Coast Ranges of southwestern Oregon and California and on the western foothills of the Sierra Nevada southward to the Transverse Ranges and the Tehachapi Mountains of Ventura and Kern counties, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 458. FNA vol. 5, p. 450.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Eriogonella
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms C. palmeri var. ventricosa Eriogonella membranacea
Name authority Goodman: Leafl. W. Bot. 2: 193, figs. 1, 2. (1939) Bentham: Trans. Linn. Soc. London 17: 419, plate 17, fig. 11. (1836)
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