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starlet spineflower, starlite spineflower

riverside spineflower, Xantus' spineflower

Habit Plants erect, 0.5–2.5(–3) × 0.5–3 dm, hirsute. Plants erect to infrequently spreading, (0.3–)0.5–2.5(–3) × 0.5–3(–5) dm, thinly pubescent.
Leaves

basal;

petiole 0.1–0.5 cm;

blade narrowly lanceolate to oblanceolate, 0.5–2 × 0.8–2(–2.2) cm, hirsute.

basal or nearly so;

petiole 1–2(–3) cm;

blade oblong or oblong-ovate to ovate, 0.3–1(–1.5) × 0.3–0.8(–1) cm, thinly pubescent adaxially, densely tomentose abaxially.

Inflorescences

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to leaves at proximal nodes only reduced, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 10–30(–40) mm, awns absent.

mostly flat-topped and openly branched, usually reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, oblong-ovate to ovate, 0.3–0.8 cm × 2–6 mm, becoming sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.4 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

Involucres

congested in small bracteated terminal clusters of 2–4 at node of dichotomies, tannish, cylindric, slightly ventricose basally, 3–4 mm, with conspicuous, white, broad, membranous margins typically extending up tooth to awn, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, 1–1.5 mm, awns straight, 0.5–1 mm.

in open clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4.5 mm, not corrugate, without scarious or membranous margins, thinly to densely pubescent;

teeth spreading, unequal, 0.7–1.5 mm, 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.5–1 mm.

Flowers

exserted;

perianth cream to creamy white or rose, cylindric, 4–4.5(–5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, obcordate to 2-lobed apically, sometimes slightly irregular but not distinctly erose;

stamens 9, slightly exserted;

filaments distinct, 4–5 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 2/3 their length, monomorphic to slightly dimorphic, narrowly oblanceolate, rounded apically, those of outer whorl occasionally slightly broader and longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

Achenes

light brown, globose-lenticular, 3.5–4.5 mm.

brown, lenticular, 4–4.5 mm.

2n

= 38, 40, 44.

Chorizanthe stellulata

Chorizanthe xanti

Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, mixed grassland and chaparral communities, oak-pine woodlands
Elevation 30-900 m (100-3000 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe stellulata can be locally common in the foothills bordering the Central Valley from Shasta County south to Stanislaus County on the western side, and to Tulare County on the eastern side. Post-flowering specimens of starlite spineflower and Douglas’s spineflower are sometimes difficult to distinguish. The margins of the involucre in the former are always white; those of C. douglasii are purple.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres thinly pubescent
var. xanti
1. Involucres densely white-pubescent
var. leucotheca
Source FNA vol. 5, p. 452. FNA vol. 5, p. 463.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri
Subordinate taxa
C. xanti var. leucotheca, C. xanti var. xanti
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle., Prodr. 14: 26. (1856) S. Watson: Proc. Amer. Acad. Arts 12: 272. (1877)
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