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Mojave spineflower

San Francisco spineflower

Habit Plants spreading to prostrate, 0.3–0.8(–1) × 0.5–8 dm. Plants decumbent to prostrate or ascending, 0.5–2(–2.5) × 0.5–10 dm, villous.
Leaves

basal;

petiole 0.5–2 cm;

blade (0.3–)0.5–1.5(–2) × (3–)5–10(–12) mm, thinly pubescent adaxially, more densely so to tomentose abaxially.

basal;

petiole (0.5–)1–3 cm;

blade oblanceolate, (0.5–)1–5 × (0.3–)0.4–0.7(–1) cm, villous.

Inflorescences

greenish to reddish, mostly flat-topped and open to dense;

bracts 3, whorled, short-petiolate, linear-lanceolate to lanceolate, acerose, 0.5–1.5 cm × 3–8(–10) mm, awns straight, 1–3.5 mm.

rather dense with secondary branches suppressed, greenish to reddish;

bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming narrowly elliptic to linear-lanceolate and aciculate at distal nodes, acerose, 0.5–5 cm × 2–7 mm, awns 0.5–1.2 mm.

Involucres

usually congested in small terminal clusters of 1–3 at node of dichotomies, (4–)5-ribbed, weakly 3-angled, 2–2.5 mm, not corrugate, densely canescent;

teeth (4–)5, essentially erect with longer, prominent, and thickened anterior one 2–4 mm, with straight awn 1–2.5 mm, remaining teeth smaller, 0.5–1 mm, with straight awns 0.3–0.8 mm.

1, greenish, cylindric, often ventricose basally, 1–3 mm, without scarious margins or if so then white to pink, thin, and restricted to basal portions of teeth, corrugate, villous abaxially;

teeth spreading, equal, 0.5–2 mm;

awns uncinate or straight with longer ones 2–3 mm and anterior one mostly 2.5–3 mm, these alternating with shorter 1–1.5(–1.7) mm ones.

Flowers

1, exserted;

perianth, cylindric, 2.5–3.5 mm;

tepals connate 1/2–2/3 their length, dimorphic, entire, those of outer whorl spreading, broadly obovate and rounded apically, those of inner whorl erect, narrowly oblanceolate, 1/2 length of outer ones, acute apically;

stamens slightly exserted;

filaments 2.5–3 mm, glabrous;

anthers yellowish, oblong, 0.5–0.7 mm.

included to slightly exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, oblong, truncate to 3-lobed and distinctly cuspidate apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, narrowly oblong, 0.3–0.4 mm.

Achenes

2.5–3 mm.

light brown, globose-lenticular, 2–3 mm.

2n

= (40), 44, (46).

Chorizanthe spinosa

Chorizanthe cuspidata

Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, saltbush communities
Elevation 600-1300 m (2000-4300 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

G. J. Goodman (1934) referred Chorizanthe spinosa to Eriogonella, but C. spinosa and C. membranacea, the type of Eriogonella, are well isolated from one another, and both are well removed from the remainder of the annual spineflowers. The Mojave spineflower is local and uncommon from southeastern Kern and southern Inyo counties, south into adjacent northeastern Los Angeles and northwestern San Bernardino counties to Antelope and Lucerne valleys.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral awns uncinate apically; San Francisco area
var. cuspidata
1. Involucral awns straight, rarely curved or with 1 awn uncinate; Marin and Sonoma counties
var. villosa
Source FNA vol. 5, p. 449. FNA vol. 5, p. 454.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Quintaria Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. cuspidata var. cuspidata, C. cuspidata var. villosa
Synonyms Eriogonella spinosa C. pungens var. cuspidata
Name authority S. Watson: in W. H. Brewer et al., Bot. California 2: 481. (1880) S. Watson: Proc. Amer. Acad. Arts 17: 379. (1882)
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