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robust spineflower, Scotts Valley spineflower

five-tooth spineflower, Watson's spineflower

Habit Plants erect to spreading or decumbent, 0.5–3 × 0.1–6 dm, villous. Plants spreading to erect, 0.2–1(–1.5) × 0.2–1(–1.5) dm, densely canescent-strigose.
Leaves

basal or nearly so;

petiole 1–4(–7) cm;

blade oblanceolate, 1.5–5 × 0.2–0.7(–1) cm, villous.

basal or nearly so;

petiole (0.5–)1–2.5(–3) cm;

blade oblanceolate, (0.3–)0.5–1.5(–2) × 0.2–0.4(–0.5) cm, thinly floccose to sparsely tomentose.

Inflorescences

with secondary branches not suppressed except in terminal clusters of involucres, green to reddish;

bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 1–5 cm × 2–5(–7) mm, awns absent.

with involucres in small clusters 0.5–1 cm diam., greenish to reddish;

bracts 2, with laminar ones oblanceolate, (0.5–)0.8–1.5(–2) cm × (1–)2–4 mm, those at distal nodes becoming sessile, reduced and scalelike, linear-lanceolate, acicular, awns slightly curved, 0.5–1 mm.

Involucres

1, greenish, cylindric, not ventricose, 2.5–4 mm, with white to pinkish, thin scarious margins restricted to basal portion of teeth, not corrugate, thinly pubescent abaxially;

teeth spreading, equal, 0.3–0.8(–1) mm;

awns uncinate with longer ones 0.7–1.3 mm and anterior one mostly 1–1.3 mm, these alternating with shorter (0.3–0.7 mm) ones.

1, green, cylindric, 5-ribbed, 3–4.5 mm, finely corrugate, pubescent;

teeth 5, erect, unequal, with leaflike, narrowly lanceolate, 2–6 mm anterior tooth, others linear, 1–2 mm;

awn uncinate, 0.4–0.8(–1) mm.

Flowers

slightly exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2.5–4 mm, pubescent abaxially;

tepals connate 1/4 their length, monomorphic, oblanceolate to narrowly oblong, usually truncate to rounded and erose or denticulate apically, occasionally distinctly cuspidate;

stamens 9, included;

filaments distinct, 2–3.5 mm, glabrous;

anthers pink to red or maroon, oblong, 0.6–0.8 mm.

1, included to slightly exserted;

perianth yellow, cylindric, 1.5–2.5 mm, thinly pubescent abaxially;

tepals connate ca. 2/3 their length, monomorphic, oblong, acute, entire apically, mostly erect;

stamens 3 or 9, slightly exserted;

filaments distinct, 0.8–1 mm, glabrous;

anthers yellow, ovate, 0.2–0.3 mm.

Achenes

light brown, globose-lenticular, 3.5–4 mm.

brown, lenticular, 2.5–3 mm.

Chorizanthe robusta

Chorizanthe watsonii

Phenology Flowering Apr–Aug.
Habitat Sandy to gravelly flats and slopes, mixed grassland, saltbush and sagebrush communities, pinyon-juniper woodlands
Elevation 300-2400 m (1000-7900 ft)
Distribution
from USDA
wc Calif
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; ID; NV; OR; UT; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe watsonii is widely distributed in the cold desert of the Great Basin and in the northern part of the warmer Mojave Desert. Plants in the northern part of the range (especially on the Palouse Prairie of south-eastern Washington) usually have three stamens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white; plants spreading or decumbent; sandy to gravelly places; Alameda and San Mateo counties s in mountains and near coast to n Monterey County
var. robusta
1. Margins of involucres rose-pink; plants erect; annual grasslands near Scotts Valley, Santa Cruz Mountains, Santa Cruz County
var. hartwegii
Source FNA vol. 5, p. 455. FNA vol. 5, p. 468.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. wheeleri, C. xanti
Subordinate taxa
C. robusta var. hartwegii, C. robusta var. robusta
Name authority Parry: Proc. Davenport Acad. Nat. Sci. 5: 176. (1889) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 199. (1870)
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