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robust spineflower, Scotts Valley spineflower

Mojave spineflower

Habit Plants erect to spreading or decumbent, 0.5–3 × 0.1–6 dm, villous. Plants spreading to prostrate, 0.3–0.8(–1) × 0.5–8 dm.
Leaves

basal or nearly so;

petiole 1–4(–7) cm;

blade oblanceolate, 1.5–5 × 0.2–0.7(–1) cm, villous.

basal;

petiole 0.5–2 cm;

blade (0.3–)0.5–1.5(–2) × (3–)5–10(–12) mm, thinly pubescent adaxially, more densely so to tomentose abaxially.

Inflorescences

with secondary branches not suppressed except in terminal clusters of involucres, green to reddish;

bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 1–5 cm × 2–5(–7) mm, awns absent.

greenish to reddish, mostly flat-topped and open to dense;

bracts 3, whorled, short-petiolate, linear-lanceolate to lanceolate, acerose, 0.5–1.5 cm × 3–8(–10) mm, awns straight, 1–3.5 mm.

Involucres

1, greenish, cylindric, not ventricose, 2.5–4 mm, with white to pinkish, thin scarious margins restricted to basal portion of teeth, not corrugate, thinly pubescent abaxially;

teeth spreading, equal, 0.3–0.8(–1) mm;

awns uncinate with longer ones 0.7–1.3 mm and anterior one mostly 1–1.3 mm, these alternating with shorter (0.3–0.7 mm) ones.

usually congested in small terminal clusters of 1–3 at node of dichotomies, (4–)5-ribbed, weakly 3-angled, 2–2.5 mm, not corrugate, densely canescent;

teeth (4–)5, essentially erect with longer, prominent, and thickened anterior one 2–4 mm, with straight awn 1–2.5 mm, remaining teeth smaller, 0.5–1 mm, with straight awns 0.3–0.8 mm.

Flowers

slightly exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2.5–4 mm, pubescent abaxially;

tepals connate 1/4 their length, monomorphic, oblanceolate to narrowly oblong, usually truncate to rounded and erose or denticulate apically, occasionally distinctly cuspidate;

stamens 9, included;

filaments distinct, 2–3.5 mm, glabrous;

anthers pink to red or maroon, oblong, 0.6–0.8 mm.

1, exserted;

perianth, cylindric, 2.5–3.5 mm;

tepals connate 1/2–2/3 their length, dimorphic, entire, those of outer whorl spreading, broadly obovate and rounded apically, those of inner whorl erect, narrowly oblanceolate, 1/2 length of outer ones, acute apically;

stamens slightly exserted;

filaments 2.5–3 mm, glabrous;

anthers yellowish, oblong, 0.5–0.7 mm.

Achenes

light brown, globose-lenticular, 3.5–4 mm.

2.5–3 mm.

2n

= (40), 44, (46).

Chorizanthe robusta

Chorizanthe spinosa

Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, saltbush communities
Elevation 600-1300 m (2000-4300 ft)
Distribution
from USDA
wc Calif
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

G. J. Goodman (1934) referred Chorizanthe spinosa to Eriogonella, but C. spinosa and C. membranacea, the type of Eriogonella, are well isolated from one another, and both are well removed from the remainder of the annual spineflowers. The Mojave spineflower is local and uncommon from southeastern Kern and southern Inyo counties, south into adjacent northeastern Los Angeles and northwestern San Bernardino counties to Antelope and Lucerne valleys.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white; plants spreading or decumbent; sandy to gravelly places; Alameda and San Mateo counties s in mountains and near coast to n Monterey County
var. robusta
1. Margins of involucres rose-pink; plants erect; annual grasslands near Scotts Valley, Santa Cruz Mountains, Santa Cruz County
var. hartwegii
Source FNA vol. 5, p. 455. FNA vol. 5, p. 449.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Quintaria
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. robusta var. hartwegii, C. robusta var. robusta
Synonyms Eriogonella spinosa
Name authority Parry: Proc. Davenport Acad. Nat. Sci. 5: 176. (1889) S. Watson: in W. H. Brewer et al., Bot. California 2: 481. (1880)
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