Chorizanthe rigida |
Chorizanthe uniaristata |
|
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devil's spineflower, rigid spineflower, rigid spiny-herb, spiny-herb |
one-awn spineflower |
|
Habit | Plants erect, 0.2–0.8(–1.5) × 0.1–0.7(–1) dm, pubescent. | Plants spreading or ascending, 0.2–0.6(–0.8) × 0.5–4(–5) dm, appressed-pubescent. |
Leaves | basal and cauline; petiole 0.5–3(–4) cm; basal blade broadly elliptic to obovate, 0.5–2.5 × (0.3–)0.5–2 cm, thinly pubescent adaxially, more densely so to tomentose abaxially; proximal cauline leaf soon deciduous, 1, blade similar to basal leaf blades only 1–2(–2.5) × 0.5–1.5 cm, mucronate to awn-tipped, awn mostly 2–4 mm; distal cauline leaf blade persistent, 1 per node, sessile, blade linear to linear-lanceolate, 0.1–1.5 × 0.05–0.15 cm, becoming hard and thornlike with age. |
basal; petiole 0.5–2 cm; blade oblanceolate, 0.5–1.5(–2) × 0.2–0.8 cm, thinly pubescent. |
Inflorescences | with involucres in dense clusters in axils of bracts, these on short shoots and each subtended by cauline leaves; bracts 2, subopposite to opposite, linear, 0.5–1(–1.2) cm × 1–2 mm, awns straight, 2–4 mm. |
with involucres in small open clusters 0.5–1.5 cm diam., greenish to grayish or reddish; bracts 2, sessile, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–5 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.4–1.2 cm × 1–2(–3) mm, awns straight, 1.5–4 mm. |
Involucres | 1, greenish, urceolate, 3-ribbed, 2–3 mm, corrugate, pubescent, rarely villous near base in some; teeth 3, unequal, with thickened anterior tooth toward base, 5–10 mm, sometimes expanding and becoming lanceolate to narrowly elliptic, others 0.5–1.2 mm; awns straight. |
3–10, grayish to reddish, urceolate, slightly ventricose basally, 2–3 mm, without scarious or membranous margins, slightly corrugate, densely grayish-pubescent; teeth widely spreading to divergent, unequal, 0.3–0.5 or 3–6 mm; awns straight or uncinate, unequal, with longer anterior one straight, 2.5–5.5 mm, others spreading, uncinate, 0.3–0.5 mm. |
Flowers | 1–2, included to slightly exserted; perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially; tepals connate ca. 2/3 their length, monomorphic, oblong, rounded, entire apically; stamens 9, slightly exserted; filaments distinct, 0.5–1 mm, glabrous; anthers yellowish, ovate, 0.2–0.3 mm. |
included or only slightly exserted; perianth bicolored with floral tube greenish white and tepals white, cylindric, 2–3 mm, sparsely pubescent; tepals connate 2/3 their length, dimorphic, linear-oblong, those of outer whorl spreading, narrowly oblong, 1.5 times longer than those of inner whorl, rounded but with minute cusp or 3 teeth apically, those of inner whorl erect to slightly spreading, acute, entire apically; stamens 3, included; filaments distinct, 1–2 mm, glabrous; anthers white, ovate, 0.4–0.5 mm. |
Achenes | brown, 3-gonous, (1.5–)1.8–2.2 mm. |
brown, globose-lenticular, 2–3 mm. |
2n | = 38, 40. |
= (78), 80, (82). |
Chorizanthe rigida |
Chorizanthe uniaristata |
|
Phenology | Flowering Feb–Jun. | Flowering Apr–Jul. |
Habitat | Sandy to gravelly or rocky flats and slopes, desert scrub | Sandy to gravelly talus or clay flats and slopes, mixed grassland and chaparral communities, pine-oak woodlands |
Elevation | -60-1900 m. (-200-6200 ft.) | 800-1900 m (2600-6200 ft) |
Distribution |
AZ; CA; NV; UT; Mexico (Baja California, Sonora)
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CA
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Discussion | Anyone with the misfortune to step bare-footed on Chorizanthe rigida after the plant has dried instantly appreciates its common name. The species is widespread on the Mojave and Sonoran deserts but only occasionally is it locally abundant or weedy. It is found also along the Lahontan Trough in western Nevada, a well-known biogeographic extension route north of the Mojave Desert (J. L. Reveal 1980). The exceedingly compact and dense inflorescences with suppressed secondary branches result in a series of leaves and bracts that subtend a closely arranged series of bracteated and involucrated flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Chorizanthe uniaristata is scattered in the Inner Coast Ranges and across the Transverse and Tehachapi ranges to the southern Sierra Nevada. One-awn spineflower is a polyploid, but whether an autopolyploid or an autoallopolyploid has not been determined. It has the smallest meiotic chromosomes observed by C. B. Hardham (1989). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 467. | FNA vol. 5, p. 460. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Acanthogonum rigidum | |
Name authority | (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870) | Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 195. (1870) |
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