Chorizanthe rigida |
Chorizanthe leptotheca |
|
---|---|---|
devil's spineflower, rigid spineflower, rigid spiny-herb, spiny-herb |
Peninsular spineflower, Ramona spineflower |
|
Habit | Plants erect, 0.2–0.8(–1.5) × 0.1–0.7(–1) dm, pubescent. | Plants erect to spreading, 0.5–3(–3.5) × 0.5–3(–5) dm, thinly pubescent. |
Leaves | basal and cauline; petiole 0.5–3(–4) cm; basal blade broadly elliptic to obovate, 0.5–2.5 × (0.3–)0.5–2 cm, thinly pubescent adaxially, more densely so to tomentose abaxially; proximal cauline leaf soon deciduous, 1, blade similar to basal leaf blades only 1–2(–2.5) × 0.5–1.5 cm, mucronate to awn-tipped, awn mostly 2–4 mm; distal cauline leaf blade persistent, 1 per node, sessile, blade linear to linear-lanceolate, 0.1–1.5 × 0.05–0.15 cm, becoming hard and thornlike with age. |
basal; petiole 1–3(–4) cm; blade oblong to oblong-ovate, 0.5–2(–3) × 0.3–0.5(–0.7) cm, thinly pubescent adaxially, usually densely tomentose adaxially. |
Inflorescences | with involucres in dense clusters in axils of bracts, these on short shoots and each subtended by cauline leaves; bracts 2, subopposite to opposite, linear, 0.5–1(–1.2) cm × 1–2 mm, awns straight, 2–4 mm. |
mostly flat-topped and openly branched, usually reddish; bracts soon deciduous, 2, occasionally leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, ovate, 0.3–0.4 cm × 2–3 mm, otherwise sessile, linear and acicular, often acerose, 0.1–0.3 cm × 0.7–1 mm, awns straight, 0.5–1 mm. |
Involucres | 1, greenish, urceolate, 3-ribbed, 2–3 mm, corrugate, pubescent, rarely villous near base in some; teeth 3, unequal, with thickened anterior tooth toward base, 5–10 mm, sometimes expanding and becoming lanceolate to narrowly elliptic, others 0.5–1.2 mm; awns straight. |
in congested clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4 mm, not corrugate, without scarious or membranous margins, thinly pubescent; teeth spreading, unequal, 0.7–1.5 mm with longer of 3 longest ones more erect than 3 other shorter and less-prominent ones, awns uncinate, 0.5–1 mm. |
Flowers | 1–2, included to slightly exserted; perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially; tepals connate ca. 2/3 their length, monomorphic, oblong, rounded, entire apically; stamens 9, slightly exserted; filaments distinct, 0.5–1 mm, glabrous; anthers yellowish, ovate, 0.2–0.3 mm. |
long-exserted; perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent; tepals connate ca. 1/2 their length, dimorphic or sometimes monomorphic, narrowly oblanceolate, apex rounded, those of outer whorl slightly broader and occasionally longer than those of inner whorl; stamens 9, mostly included; filaments distinct, 4–6 mm, glabrous; anthers pink to red, ovate to oblong, 0.5–0.6 mm. |
Achenes | brown, 3-gonous, (1.5–)1.8–2.2 mm. |
brown, lenticular, 3–4 mm. |
2n | = 38, 40. |
= 38. |
Chorizanthe rigida |
Chorizanthe leptotheca |
|
Phenology | Flowering Feb–Jun. | Flowering May–Aug. |
Habitat | Sandy to gravelly or rocky flats and slopes, desert scrub | Sandy to gravelly flats and slopes, grassland and chaparral communities, pine-oak woodlands |
Elevation | -60-1900 m. (-200-6200 ft.) | (300-)600-1600(-1900) m ((1000-)2000-5200(-6200) ft) |
Distribution |
AZ; CA; NV; UT; Mexico (Baja California, Sonora)
|
CA; Mexico (Baja California)
|
Discussion | Anyone with the misfortune to step bare-footed on Chorizanthe rigida after the plant has dried instantly appreciates its common name. The species is widespread on the Mojave and Sonoran deserts but only occasionally is it locally abundant or weedy. It is found also along the Lahontan Trough in western Nevada, a well-known biogeographic extension route north of the Mojave Desert (J. L. Reveal 1980). The exceedingly compact and dense inflorescences with suppressed secondary branches result in a series of leaves and bracts that subtend a closely arranged series of bracteated and involucrated flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Chorizanthe leptotheca is found in the foothills of the San Bernardino Mountains of San Bernardino County southward along the eastern edge of the Santa Ana Mountains, and through the San Jacinto and Santa Rosa mountains of Riverside County into the mountains of central San Diego County. The species is also found in north-central Baja California. Ramona spineflower is clearly related to Chorizanthe staticoides, but that species occurs to the west of the range of C. leptotheca and the two are not known to be sympatric. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 467. | FNA vol. 5, p. 463. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Acanthogonum rigidum | |
Name authority | (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870) | Goodman: Ann. Missouri Bot. Gard. 21: 61. (1934) |
Web links |