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devil's spineflower, rigid spineflower, rigid spiny-herb, spiny-herb

Douglas' spineflower, San Benito spineflower

Habit Plants erect, 0.2–0.8(–1.5) × 0.1–0.7(–1) dm, pubescent. Plants erect, 1–4(–5) × 0.5–3 dm, villous.
Leaves

basal and cauline;

petiole 0.5–3(–4) cm;

basal blade broadly elliptic to obovate, 0.5–2.5 × (0.3–)0.5–2 cm, thinly pubescent adaxially, more densely so to tomentose abaxially;

proximal cauline leaf soon deciduous, 1, blade similar to basal leaf blades only 1–2(–2.5) × 0.5–1.5 cm, mucronate to awn-tipped, awn mostly 2–4 mm;

distal cauline leaf blade persistent, 1 per node, sessile, blade linear to linear-lanceolate, 0.1–1.5 × 0.05–0.15 cm, becoming hard and thornlike with age.

basal;

petiole 1–3(–6) cm;

blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous.

Inflorescences

with involucres in dense clusters in axils of bracts, these on short shoots and each subtended by cauline leaves;

bracts 2, subopposite to opposite, linear, 0.5–1(–1.2) cm × 1–2 mm, awns straight, 2–4 mm.

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent.

Involucres

1, greenish, urceolate, 3-ribbed, 2–3 mm, corrugate, pubescent, rarely villous near base in some;

teeth 3, unequal, with thickened anterior tooth toward base, 5–10 mm, sometimes expanding and becoming lanceolate to narrowly elliptic, others 0.5–1.2 mm;

awns straight.

congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm.

Flowers

1–2, included to slightly exserted;

perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially;

tepals connate ca. 2/3 their length, monomorphic, oblong, rounded, entire apically;

stamens 9, slightly exserted;

filaments distinct, 0.5–1 mm, glabrous;

anthers yellowish, ovate, 0.2–0.3 mm.

exserted;

perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire;

stamens 9, slightly exserted;

filaments distinct, 3–4 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

Achenes

brown, 3-gonous, (1.5–)1.8–2.2 mm.

light brown, globose-lenticular, 3.5–4 mm.

2n

= 38, 40.

= 40.

Chorizanthe rigida

Chorizanthe douglasii

Phenology Flowering Feb–Jun. Flowering Apr–Jul.
Habitat Sandy to gravelly or rocky flats and slopes, desert scrub Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands
Elevation -60-1900 m. (-200-6200 ft.) (200-)300-1600 m ((700-)1000-5200 ft)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Anyone with the misfortune to step bare-footed on Chorizanthe rigida after the plant has dried instantly appreciates its common name. The species is widespread on the Mojave and Sonoran deserts but only occasionally is it locally abundant or weedy. It is found also along the Lahontan Trough in western Nevada, a well-known biogeographic extension route north of the Mojave Desert (J. L. Reveal 1980). The exceedingly compact and dense inflorescences with suppressed secondary branches result in a series of leaves and bracts that subtend a closely arranged series of bracteated and involucrated flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 467. FNA vol. 5, p. 452.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms Acanthogonum rigidum C. nortonii
Name authority (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870) Bentham: Trans. Linn. Soc. London 17: 418. (1836)
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