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devil's spineflower, rigid spineflower, rigid spiny-herb, spiny-herb

Cleveland's spineflower

Habit Plants erect, 0.2–0.8(–1.5) × 0.1–0.7(–1) dm, pubescent. Plants spreading to decumbent, 0.2–0.8(–1) × 0.5–5(–7) dm, appressed-pubescent.
Leaves

basal and cauline;

petiole 0.5–3(–4) cm;

basal blade broadly elliptic to obovate, 0.5–2.5 × (0.3–)0.5–2 cm, thinly pubescent adaxially, more densely so to tomentose abaxially;

proximal cauline leaf soon deciduous, 1, blade similar to basal leaf blades only 1–2(–2.5) × 0.5–1.5 cm, mucronate to awn-tipped, awn mostly 2–4 mm;

distal cauline leaf blade persistent, 1 per node, sessile, blade linear to linear-lanceolate, 0.1–1.5 × 0.05–0.15 cm, becoming hard and thornlike with age.

basal;

petiole 0.5–2 mm;

blade oblanceolate, 0.5–1.5(–2) × 0.3–0.6(–0.8) cm, thinly pubescent.

Inflorescences

with involucres in dense clusters in axils of bracts, these on short shoots and each subtended by cauline leaves;

bracts 2, subopposite to opposite, linear, 0.5–1(–1.2) cm × 1–2 mm, awns straight, 2–4 mm.

with involucres in small, open clusters 0.5–1.5 cm diam., greenish or grayish to reddish;

bracts 2, sessile, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–5 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.4–1 cm × 1–2(–3) mm, awns straight, 1–3 mm.

Involucres

1, greenish, urceolate, 3-ribbed, 2–3 mm, corrugate, pubescent, rarely villous near base in some;

teeth 3, unequal, with thickened anterior tooth toward base, 5–10 mm, sometimes expanding and becoming lanceolate to narrowly elliptic, others 0.5–1.2 mm;

awns straight.

3–10, grayish to reddish, urceolate, slightly ventricose basally, 3–3.5 mm, slightly corrugate, without scarious or membranous margins, densely pubescent;

teeth widely spreading to divergent, unequal, 0.3–0.6 mm or 3–6 mm;

awns uncinate, unequal, with longer anterior one 1.5–2.5 mm, others spreading, 0.3–0.6 mm.

Flowers

1–2, included to slightly exserted;

perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially;

tepals connate ca. 2/3 their length, monomorphic, oblong, rounded, entire apically;

stamens 9, slightly exserted;

filaments distinct, 0.5–1 mm, glabrous;

anthers yellowish, ovate, 0.2–0.3 mm.

included or only slightly exserted;

perianth bicolored with floral tube greenish white and tepals white, cylindric, 2.5–3 mm, sparsely pubescent;

tepals connate 2/3 their length, dimorphic, linear-oblong, those of outer whorl spreading, 1.5 times longer than those of inner whorl, rounded, entire or emarginate to slightly 2-lobed apically, those of inner whorl erect, acute, entire to erose, slightly fimbriate or 2-lobed apically;

stamens 3, included;

filaments distinct, 2–2.5 mm, glabrous;

anthers white, ovate, 0.3–0.4 mm.

Achenes

brown, 3-gonous, (1.5–)1.8–2.2 mm.

brown, globose-lenticular, 2.5–3 mm.

2n

= 38, 40.

= 42.

Chorizanthe rigida

Chorizanthe clevelandii

Phenology Flowering Feb–Jun. Flowering May–Sep.
Habitat Sandy to gravelly or rocky flats and slopes, desert scrub Sandy to gravelly flats and slopes, mixed grassland and chaparral communities, pine-oak woodlands
Elevation -60-1900 m. (-200-6200 ft.) 400-2000 m (1300-6600 ft)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Anyone with the misfortune to step bare-footed on Chorizanthe rigida after the plant has dried instantly appreciates its common name. The species is widespread on the Mojave and Sonoran deserts but only occasionally is it locally abundant or weedy. It is found also along the Lahontan Trough in western Nevada, a well-known biogeographic extension route north of the Mojave Desert (J. L. Reveal 1980). The exceedingly compact and dense inflorescences with suppressed secondary branches result in a series of leaves and bracts that subtend a closely arranged series of bracteated and involucrated flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe clevelandii is locally infrequent to common in scattered locations in the Coast Ranges from Mendocino and Lake counties south to Santa Barbara County, and across the Transverse and Tehachapi ranges of Ventura and Kern counties to the southern Sierra Nevada in Tulare County. It is the most widely distributed of the spineflowers endemic to California. The involucres stick to fur, clothing, and fingers, aiding dispersal of the achenes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 467. FNA vol. 5, p. 460.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms Acanthogonum rigidum
Name authority (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870) Parry: Proc. Davenport Acad. Nat. Sci. 4: 62. (1884)
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