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devil's spineflower, rigid spineflower, rigid spiny-herb, spiny-herb

Brewer's spineflower, San Luis obispo spineflower

Habit Plants erect, 0.2–0.8(–1.5) × 0.1–0.7(–1) dm, pubescent. Plants ascending to decumbent, (0.3–)0.5–1.5(–2) × 1–5(–7) dm, thinly pubescent.
Leaves

basal and cauline;

petiole 0.5–3(–4) cm;

basal blade broadly elliptic to obovate, 0.5–2.5 × (0.3–)0.5–2 cm, thinly pubescent adaxially, more densely so to tomentose abaxially;

proximal cauline leaf soon deciduous, 1, blade similar to basal leaf blades only 1–2(–2.5) × 0.5–1.5 cm, mucronate to awn-tipped, awn mostly 2–4 mm;

distal cauline leaf blade persistent, 1 per node, sessile, blade linear to linear-lanceolate, 0.1–1.5 × 0.05–0.15 cm, becoming hard and thornlike with age.

basal;

petiole 1–3 cm;

blade spatulate to ovate, 0.5–2 × 0.3–1.2(–1.5) cm, thinly pubescent to densely tomentose at least abaxially.

Inflorescences

with involucres in dense clusters in axils of bracts, these on short shoots and each subtended by cauline leaves;

bracts 2, subopposite to opposite, linear, 0.5–1(–1.2) cm × 1–2 mm, awns straight, 2–4 mm.

mostly flat-topped, open to rather densely branched, greenish to reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to leaf blades, short-petiolate, oblong to ovate, 0.3–0.8 cm × 2–6 mm, becoming sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

Involucres

1, greenish, urceolate, 3-ribbed, 2–3 mm, corrugate, pubescent, rarely villous near base in some;

teeth 3, unequal, with thickened anterior tooth toward base, 5–10 mm, sometimes expanding and becoming lanceolate to narrowly elliptic, others 0.5–1.2 mm;

awns straight.

in open clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 2.5–3 mm, corrugate, without scarious or membranous margins, thinly pubescent with slender curly hairs;

teeth spreading, unequal, 0.4–1.2 mm with 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.3–0.6 mm.

Flowers

1–2, included to slightly exserted;

perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially;

tepals connate ca. 2/3 their length, monomorphic, oblong, rounded, entire apically;

stamens 9, slightly exserted;

filaments distinct, 0.5–1 mm, glabrous;

anthers yellowish, ovate, 0.2–0.3 mm.

exserted;

perianth white to rose or red, usually with white lobes, cylindric, 3–3.5 mm, pubescent;

tepals connate ca. 1/2 their length, monomorphic to slightly dimorphic, narrowly oblong to obovate, rounded apically, those of outer whorl usually slightly broader and longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 2.5–3 mm, glabrous;

anthers pink to red, oblong, 0.4–0.5 mm.

Achenes

brown, 3-gonous, (1.5–)1.8–2.2 mm.

brown, lenticular, 2.5–3 mm.

2n

= 38, 40.

= 38.

Chorizanthe rigida

Chorizanthe breweri

Phenology Flowering Feb–Jun. Flowering Mar–Jul.
Habitat Sandy to gravelly or rocky flats and slopes, desert scrub Gravelly or rocky places, serpentine outcrops, mixed grassland and chaparral communities, pine-oak woodlands
Elevation -60-1900 m. (-200-6200 ft.) 60-800 m (200-2600 ft)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Anyone with the misfortune to step bare-footed on Chorizanthe rigida after the plant has dried instantly appreciates its common name. The species is widespread on the Mojave and Sonoran deserts but only occasionally is it locally abundant or weedy. It is found also along the Lahontan Trough in western Nevada, a well-known biogeographic extension route north of the Mojave Desert (J. L. Reveal 1980). The exceedingly compact and dense inflorescences with suppressed secondary branches result in a series of leaves and bracts that subtend a closely arranged series of bracteated and involucrated flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chorizanthe breweri is found infrequently in the Coast Ranges of southwestern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 467. FNA vol. 5, p. 464.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms Acanthogonum rigidum
Name authority (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870) S. Watson: Proc. Amer. Acad. Arts 12: 270. (1877)
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