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prickly spineflower, straight-awn spineflower

Santa Barbara spineflower, Wheeler's spineflower

Habit Plants spreading to decumbent, 0.3–0.8(–1) × 0.5–4(–5) dm, appressed-pubescent. Plants erect to spreading, 0.5–2(–2.5) × 1–2 dm, thinly pubescent.
Leaves

basal;

petiole 0.5–2 cm;

blade oblanceolate to spatulate, 0.5–1.5(–2) × 0.2–0.6 cm, thinly pubescent.

basal;

petiole 0.5–3 cm;

blade elliptic to oblong, 0.5–2 × 0.2–0.6 cm, thinly pubescent adaxially, tomentose abaxially.

Inflorescences

with involucres in small, open clusters 0.5–1.5 cm diam., greenish to grayish;

bracts 2, without whorl of sessile bracts about midstem, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–5 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.3–0.8 cm × 1–2 mm, awns straight, 0.5–1.5 mm.

mostly flat-topped, openly branched, greenish to reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to leaf blades, short-petiolate, oblong, 0.5–1.2 cm × 2–4 mm, sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

Involucres

3–10+, grayish to reddish, urceolate, slightly ventricose basally, 2–2.5(–3) mm, slightly corrugate, without scarious or membranous margins, densely pubescent;

teeth spreading, unequal, 1–2 mm;

awns straight or uncinate, unequal, with longer anterior one straight, mostly 1.5–2.5 mm, others uncinate, 0.3–0.6 mm.

in dense terminal clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 2–2.5 mm, corrugate, without scarious or membranous margins, thinly pubescent with stoutish, recurved hairs;

teeth spreading, unequal, 0.3–0.8(–1) mm, with 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.3–0.5 mm.

Flowers

exserted;

perianth bicolored with floral tube yellow and tepals yellow or white, cylindric, 3.5–4 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, obovate, those of outer whorl white, obovate to nearly orbiculate, 3–4 times longer than those of inner whorl, , truncate to slightly 2-lobed apically, those of inner lobes erect, yellow, broadly obovate, truncate and erose apically;

stamens 9, included;

filaments distinct, 1–1.5 mm, glabrous;

anthers yellow to golden, oblong, 0.5–0.6 mm.

exserted;

perianth white or rose or red with white lobes, cylindric, 2.5–3 mm, glabrous except for few scattered hairs ca. midlength along midrib abaxially;

tepals connate 1/2 their length, monomorphic to slightly dimorphic, oblong, rounded apically, those of outer whorl usually slightly broader and longer than those of inner whorl;

stamens 6, included;

filaments distinct, 1.5–2 mm, glabrous;

anthers pink to red, oblong, 0.3–0.4 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

brown, lenticular, 2.5–3 mm.

2n

= (36), 40, (44).

Chorizanthe rectispina

Chorizanthe wheeleri

Phenology Flowering May–Jul. Flowering Apr–Jun.
Habitat Sandy to gravelly flats and slopes, mixed grassland communities, pine-oak woodlands Gravelly to rocky slopes, coastal scrub communities
Elevation 200-600 m (700-2000 ft) 0-400(-600) m (0-1300(-2000) ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chorizanthe rectispina is infrequent and localized in the Coast Ranges of west-central California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe wheeleri is a rare insular endemic known only from Santa Cruz and Santa Rosa islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 460. FNA vol. 5, p. 464.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. xanti
Synonyms C. insularis
Name authority Goodman: Ann. Missouri Bot. Gard. 21: 72. (1934) S. Watson: Proc. Amer. Acad. Arts 12: 272. (1877)
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