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prickly spineflower, straight-awn spineflower

San Francisco spineflower

Habit Plants spreading to decumbent, 0.3–0.8(–1) × 0.5–4(–5) dm, appressed-pubescent. Plants decumbent to prostrate or ascending, 0.5–2(–2.5) × 0.5–10 dm, villous.
Leaves

basal;

petiole 0.5–2 cm;

blade oblanceolate to spatulate, 0.5–1.5(–2) × 0.2–0.6 cm, thinly pubescent.

basal;

petiole (0.5–)1–3 cm;

blade oblanceolate, (0.5–)1–5 × (0.3–)0.4–0.7(–1) cm, villous.

Inflorescences

with involucres in small, open clusters 0.5–1.5 cm diam., greenish to grayish;

bracts 2, without whorl of sessile bracts about midstem, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–5 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.3–0.8 cm × 1–2 mm, awns straight, 0.5–1.5 mm.

rather dense with secondary branches suppressed, greenish to reddish;

bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming narrowly elliptic to linear-lanceolate and aciculate at distal nodes, acerose, 0.5–5 cm × 2–7 mm, awns 0.5–1.2 mm.

Involucres

3–10+, grayish to reddish, urceolate, slightly ventricose basally, 2–2.5(–3) mm, slightly corrugate, without scarious or membranous margins, densely pubescent;

teeth spreading, unequal, 1–2 mm;

awns straight or uncinate, unequal, with longer anterior one straight, mostly 1.5–2.5 mm, others uncinate, 0.3–0.6 mm.

1, greenish, cylindric, often ventricose basally, 1–3 mm, without scarious margins or if so then white to pink, thin, and restricted to basal portions of teeth, corrugate, villous abaxially;

teeth spreading, equal, 0.5–2 mm;

awns uncinate or straight with longer ones 2–3 mm and anterior one mostly 2.5–3 mm, these alternating with shorter 1–1.5(–1.7) mm ones.

Flowers

exserted;

perianth bicolored with floral tube yellow and tepals yellow or white, cylindric, 3.5–4 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, obovate, those of outer whorl white, obovate to nearly orbiculate, 3–4 times longer than those of inner whorl, , truncate to slightly 2-lobed apically, those of inner lobes erect, yellow, broadly obovate, truncate and erose apically;

stamens 9, included;

filaments distinct, 1–1.5 mm, glabrous;

anthers yellow to golden, oblong, 0.5–0.6 mm.

included to slightly exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, oblong, truncate to 3-lobed and distinctly cuspidate apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, narrowly oblong, 0.3–0.4 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

light brown, globose-lenticular, 2–3 mm.

2n

= (36), 40, (44).

Chorizanthe rectispina

Chorizanthe cuspidata

Phenology Flowering May–Jul.
Habitat Sandy to gravelly flats and slopes, mixed grassland communities, pine-oak woodlands
Elevation 200-600 m (700-2000 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chorizanthe rectispina is infrequent and localized in the Coast Ranges of west-central California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral awns uncinate apically; San Francisco area
var. cuspidata
1. Involucral awns straight, rarely curved or with 1 awn uncinate; Marin and Sonoma counties
var. villosa
Source FNA vol. 5, p. 460. FNA vol. 5, p. 454.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. cuspidata var. cuspidata, C. cuspidata var. villosa
Synonyms C. pungens var. cuspidata
Name authority Goodman: Ann. Missouri Bot. Gard. 21: 72. (1934) S. Watson: Proc. Amer. Acad. Arts 17: 379. (1882)
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