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Monterey spineflower

pink spineflower

Habit Plants prostrate to ascending or erect, 0.5–2(–2.5) × 0.5–10 dm, grayish-villous. Plants 1–6(–10) × 0.5–3(–5) dm, woolly-floccose.
Leaves

basal;

petiole (0.5–)1–3(–4) cm;

blade oblanceolate, (0.5–)1–5(–7) × (0.3–)0.4–0.7(–1) cm, villous.

basal and cauline;

petiole 0.1–0.5(–0.8) cm;

blade linear to narrowly oblanceolate, (1–)1.5–5 × 0.1–0.3 cm, thinly to densely floccose adaxially, densely tomentose abaxially.

Inflorescences

rather dense with secondary branches suppressed, grayish;

bracts 2, similar to leaf blades at proximal nodes only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–7 cm × 2–7 mm, awns 0.5–1.2 mm.

strict, white to greenish, open;

bracts usually 2, opposite, rarely in whorls of 3–5, short-petiolate, acerose, similar to proximal leaf blades only reduced, 0.3–3 cm × 1–3 mm, awns straight, 0.5–1 mm.

Involucres

1, grayish, cylindric, often ventricose basally, 2–2.5(–3) mm, with distinct, white to pink or purple, scarious margins extending nearly full length of awn, corrugate, villous abaxially;

teeth spreading, equal, 0.5–1.5 mm;

awns uncinate with longer ones 2–3 mm and alternating with shorter (1–1.5 mm) ones.

usually congested in small terminal clusters of 1–3 at node of dichotomies, urceolate, ventricose basally, 3-angled, 6-ribbed, 3–4 mm, not corrugate, with conspicuous, white margins extending across sinuses, tomentose to floccose or glabrate with age, greenish to brownish;

teeth 6;

awns uncinate, 0.7–1.5 mm.

Flowers

exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3.5 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, obovate to oblong, acute to truncate and erose apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, ovate, 0.3–0.4 mm.

1(–2), slightly exserted;

perianth white to rose, subcylindric, (1.5–)2.5–3 mm, densely pubescent abaxially;

tepals connate 2/3 their length, slightly dimorphic, entire and rounded apically, those of outer whorl obovate, those of inner whorl spatulate;

stamens slightly exserted;

filaments 1.5–2.5 mm, glabrous;

anthers pink to red, oval, 0.2–0.3 mm.

Achenes

dark brown, globose-lenticular, 2–2.5 mm.

2.5–3 mm.

2n

= 38, 40, (42), 80, 82, 84.

Chorizanthe pungens

Chorizanthe membranacea

Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly or rocky flats and slopes, mixed grassland and chaparral communities, oak-pine woodlands
Elevation 40-1400(-1600) m (100-4600(-5200) ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe membranacea has long been considered an isolated element among the spineflowers. The strict, upright habit, numerous basal and cauline leaves, and broad, continuous, membranous margins of the involucre all reflect that isolation. Pink spineflower is widespread and often locally common in the Coast Ranges of southwestern Oregon and California and on the western foothills of the Sierra Nevada southward to the Transverse Ranges and the Tehachapi Mountains of Ventura and Kern counties, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white (rarely pinkish), scarious; plants prostrate to slightly ascending; coastal areas and adjacent inland valleys
var. pungens
1. Margins of involucres dark pinkish to purple, scarious; plants slightly ascending to erect; coastal mountains
var. hartwegiana
Source FNA vol. 5, p. 453. FNA vol. 5, p. 450.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Eriogonella
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. pungens var. hartwegiana, C. pungens var. pungens
Synonyms Eriogonella membranacea
Name authority Bentham: Trans. Linn. Soc. London 17: 419, plate 19, fig. 2. (1836) Bentham: Trans. Linn. Soc. London 17: 419, plate 17, fig. 11. (1836)
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