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Monterey spineflower

Peninsular spineflower, Ramona spineflower

Habit Plants prostrate to ascending or erect, 0.5–2(–2.5) × 0.5–10 dm, grayish-villous. Plants erect to spreading, 0.5–3(–3.5) × 0.5–3(–5) dm, thinly pubescent.
Leaves

basal;

petiole (0.5–)1–3(–4) cm;

blade oblanceolate, (0.5–)1–5(–7) × (0.3–)0.4–0.7(–1) cm, villous.

basal;

petiole 1–3(–4) cm;

blade oblong to oblong-ovate, 0.5–2(–3) × 0.3–0.5(–0.7) cm, thinly pubescent adaxially, usually densely tomentose adaxially.

Inflorescences

rather dense with secondary branches suppressed, grayish;

bracts 2, similar to leaf blades at proximal nodes only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–7 cm × 2–7 mm, awns 0.5–1.2 mm.

mostly flat-topped and openly branched, usually reddish;

bracts soon deciduous, 2, occasionally leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, ovate, 0.3–0.4 cm × 2–3 mm, otherwise sessile, linear and acicular, often acerose, 0.1–0.3 cm × 0.7–1 mm, awns straight, 0.5–1 mm.

Involucres

1, grayish, cylindric, often ventricose basally, 2–2.5(–3) mm, with distinct, white to pink or purple, scarious margins extending nearly full length of awn, corrugate, villous abaxially;

teeth spreading, equal, 0.5–1.5 mm;

awns uncinate with longer ones 2–3 mm and alternating with shorter (1–1.5 mm) ones.

in congested clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4 mm, not corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 0.7–1.5 mm with longer of 3 longest ones more erect than 3 other shorter and less-prominent ones, awns uncinate, 0.5–1 mm.

Flowers

exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3.5 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, obovate to oblong, acute to truncate and erose apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, ovate, 0.3–0.4 mm.

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 1/2 their length, dimorphic or sometimes monomorphic, narrowly oblanceolate, apex rounded, those of outer whorl slightly broader and occasionally longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, ovate to oblong, 0.5–0.6 mm.

Achenes

dark brown, globose-lenticular, 2–2.5 mm.

brown, lenticular, 3–4 mm.

2n

= 38.

Chorizanthe pungens

Chorizanthe leptotheca

Phenology Flowering May–Aug.
Habitat Sandy to gravelly flats and slopes, grassland and chaparral communities, pine-oak woodlands
Elevation (300-)600-1600(-1900) m ((1000-)2000-5200(-6200) ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe leptotheca is found in the foothills of the San Bernardino Mountains of San Bernardino County southward along the eastern edge of the Santa Ana Mountains, and through the San Jacinto and Santa Rosa mountains of Riverside County into the mountains of central San Diego County. The species is also found in north-central Baja California.

Ramona spineflower is clearly related to Chorizanthe staticoides, but that species occurs to the west of the range of C. leptotheca and the two are not known to be sympatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white (rarely pinkish), scarious; plants prostrate to slightly ascending; coastal areas and adjacent inland valleys
var. pungens
1. Margins of involucres dark pinkish to purple, scarious; plants slightly ascending to erect; coastal mountains
var. hartwegiana
Source FNA vol. 5, p. 453. FNA vol. 5, p. 463.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. pungens var. hartwegiana, C. pungens var. pungens
Name authority Bentham: Trans. Linn. Soc. London 17: 419, plate 19, fig. 2. (1836) Goodman: Ann. Missouri Bot. Gard. 21: 61. (1934)
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