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Orcutt spineflower, Orcutt's spineflower, San Diego spineflower

Parry's spineflower, San Bernardino spineflower, San Fernando Valley spineflower

Habit Plants prostrate, 0.1–0.5 × 0.3–2(–2.5) dm, villous. Plants prostrate to spreading, 0.2–0.8(–1) × 0.5–4(–6) dm, strigose.
Leaves

basal;

petiole 1–2 cm;

blade narrowly oblanceolate, 0.5–1.5 × 0.2–0.35(–0.5) cm, thinly pubescent.

basal;

petiole 0.5–2(–3.5) cm;

blade oblanceolate to narrowly oblong, 0.5–2.5(–4) × 0.2–0.6(–1.2) cm, thinly pubescent.

Inflorescences

with involucres in small clusters 0.5–1 cm diam., greenish;

bracts 2, sessile, unequal, 1 laminar and oblanceolate, 0.3–1 cm × 1–3 mm, awnless, this opposite linear, acicular, greatly reduced, 0.1–0.2 cm × 0.3–0.6 mm bract terminated by short, straight awn 0.6–1 mm.

with involucres in small, open clusters 0.3–1 cm diam., greenish or grayish to reddish;

bracts 2, sessile, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–7 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.1–0.5 cm × 1–2 mm, awns straight, 0.4–1 mm.

Involucres

1, greenish, campanulate, 3-ribbed, 0.8–2 mm, faintly corrugate, pubescent;

teeth 3, equal, 1.8–2 mm;

awns uncinate, 0.6–1 mm.

3–5, greenish or grayish to reddish, urceolate, slightly ventricose basally, 1.5–2 mm, corrugate, without scarious or membranous margins, pubescent;

teeth widely spreading to divergent or recurved, equal, 0.5–1.5 mm or 1–3 mm;

awns uncinate or straight, unequal, alternating 0.5–1.5 mm and 0.2–0.5 mm.

Flowers

1, included to slightly exserted;

perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially;

tepals connate ca. 1/2 their length, monomorphic, narrowly oblanceolate, obtuse to truncate, entire apically, slightly spreading;

stamens 9, slightly exserted;

filaments distinct, 0.5–0.8 mm, glabrous;

anthers reddish, ovate, 0.2–0.3 mm.

slightly exserted;

perianth bicolored with floral tube greenish white and tepals white, cylindric, 2.5–3 mm, sparsely pubescent;

tepals connate 2/3 their length, slightly dimorphic, those of outer whorl oblong to oblong-ovate, 1.5 times longer than those of inner whorl, rounded, erose or rarely some entire to denticulate apically, those of inner whorl linear-oblanceolate, acute, entire or denticulate apically;

stamens 9, included;

filaments distinct, 2–2.5 mm, glabrous;

anthers white, ovate, 0.2–0.3 mm.

Achenes

dark brown, lenticular, 2–2.2 mm.

brown, globose-lenticular, 2.5–3 mm.

2n

= (76, 78), 80, (84).

Chorizanthe orcuttiana

Chorizanthe parryi

Phenology Flowering Mar–May.
Habitat Sandy soil, mesas and hills near coast, coastal scrub communities
Elevation 60-200 m (200-700 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chorizanthe orcuttiana is known from a few populations on coastal mesas and hills near San Diego, San Diego County. It is federally listed as endangered. The species is an octoploid that may well have resulted from an ancient hybridization and doubling of chromosomes involving C. procumbens and C. polygonoides var. longispina. The Orcutt spineflower grows in soft, white sand; C. procumbens and C. polygonoides var. longispina are restricted to gravelly sites.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral awns uncinate
var. parryi
1. Involucral awns straight
var. fernandina
Source FNA vol. 5, p. 467. FNA vol. 5, p. 461.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. parryi var. fernandina, C. parryi var. parryi
Name authority Parry: Proc. Davenport Acad. Nat. Sci. 4: 54. (1884) S. Watson: Proc. Amer. Acad. Arts 12: 271. (1877)
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