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Orcutt spineflower, Orcutt's spineflower, San Diego spineflower

Palmer's spineflower

Habit Plants prostrate, 0.1–0.5 × 0.3–2(–2.5) dm, villous. Plants erect to spreading, (0.5–)1–3(–4) × 1–3 dm, appressed-pubescent.
Leaves

basal;

petiole 1–2 cm;

blade narrowly oblanceolate, 0.5–1.5 × 0.2–0.35(–0.5) cm, thinly pubescent.

basal;

petiole 1–3 cm;

blade oblanceolate, 1–3 × 0.4–0.8 cm, thinly pubescent.

Inflorescences

with involucres in small clusters 0.5–1 cm diam., greenish;

bracts 2, sessile, unequal, 1 laminar and oblanceolate, 0.3–1 cm × 1–3 mm, awnless, this opposite linear, acicular, greatly reduced, 0.1–0.2 cm × 0.3–0.6 mm bract terminated by short, straight awn 0.6–1 mm.

with involucres in dense clusters 2–4 cm diam., greenish or reddish to purple;

bracts 2–3 at proximal node, usually leaflike, often with whorl of sessile bracts about midstem, elliptic, 0.5–1.5 cm × 2–6 mm, becoming gradually lanceolate to elliptic, 0.2–1 cm × 1.5–5 mm, at distal nodes scalelike, linear and aciculate, acerose, awns straight, 1–3 mm.

Involucres

1, greenish, campanulate, 3-ribbed, 0.8–2 mm, faintly corrugate, pubescent;

teeth 3, equal, 1.8–2 mm;

awns uncinate, 0.6–1 mm.

3–10 or more, reddish to purplish, urceolate, slightly ventricose basally, 3.5–4 mm, without scarious or membranous margins, slightly corrugate, thinly pubescent with slender, curly hairs;

teeth erect to spreading, unequal, 1–2 mm;

awns uncinate, 0.5–1 mm with longer anterior one mostly 1 mm.

Flowers

1, included to slightly exserted;

perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially;

tepals connate ca. 1/2 their length, monomorphic, narrowly oblanceolate, obtuse to truncate, entire apically, slightly spreading;

stamens 9, slightly exserted;

filaments distinct, 0.5–0.8 mm, glabrous;

anthers reddish, ovate, 0.2–0.3 mm.

exserted;

perianth bicolored with floral tube white to yellow and tepals red, maroon, or dark purple, cylindric, 4–5 mm, glabrous or with few scattered hairs along midrib ca. midlength;

tepals erect, connate 1/2 their length, dimorphic, obovate, those of outer whorl slightly longer than inner whorl, entire, rounded apically, those of inner whorl fimbriate and truncate or somewhat 2-lobed;

stamens 9, exserted;

filaments distinct, 4–5 mm, glabrous;

anthers pink to red or maroon, oblong, 0.9–1 mm.

Achenes

dark brown, lenticular, 2–2.2 mm.

brown, globose-lenticular, 3–3.5 mm.

2n

= (76, 78), 80, (84).

= 38, 40, (48).

Chorizanthe orcuttiana

Chorizanthe palmeri

Phenology Flowering Mar–May. Flowering May–Aug.
Habitat Sandy soil, mesas and hills near coast, coastal scrub communities Gravelly to rocky serpentine and serpentinized igneous outcrops, mixed grassland communities, pine-oak woodlands
Elevation 60-200 m (200-700 ft) 60-700 m (200-2300 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chorizanthe orcuttiana is known from a few populations on coastal mesas and hills near San Diego, San Diego County. It is federally listed as endangered. The species is an octoploid that may well have resulted from an ancient hybridization and doubling of chromosomes involving C. procumbens and C. polygonoides var. longispina. The Orcutt spineflower grows in soft, white sand; C. procumbens and C. polygonoides var. longispina are restricted to gravelly sites.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe palmeri is known only from the Santa Lucia Mountains, the San Luis Range, and the Huasna area. Populations differ slightly both morphologically and ecologically, but recognition of variants is not suggested. At full anthesis, the reddish stems, involucres, and tepals, plus the localized concentrations of individuals provide for splashes of purplish red on the otherwise grass-brown slopes. Set against the often blackish green of serpentine barrens, the plants can be spotted even from a high-flying aircraft! This species would make an excellent addition to the garden border.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 467. FNA vol. 5, p. 457.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Name authority Parry: Proc. Davenport Acad. Nat. Sci. 4: 54. (1884) S. Watson: Proc. Amer. Acad. Arts 12: 271. (1877)
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