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Peninsular spineflower, Ramona spineflower

Mojave spineflower

Habit Plants erect to spreading, 0.5–3(–3.5) × 0.5–3(–5) dm, thinly pubescent. Plants spreading to prostrate, 0.3–0.8(–1) × 0.5–8 dm.
Leaves

basal;

petiole 1–3(–4) cm;

blade oblong to oblong-ovate, 0.5–2(–3) × 0.3–0.5(–0.7) cm, thinly pubescent adaxially, usually densely tomentose adaxially.

basal;

petiole 0.5–2 cm;

blade (0.3–)0.5–1.5(–2) × (3–)5–10(–12) mm, thinly pubescent adaxially, more densely so to tomentose abaxially.

Inflorescences

mostly flat-topped and openly branched, usually reddish;

bracts soon deciduous, 2, occasionally leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, ovate, 0.3–0.4 cm × 2–3 mm, otherwise sessile, linear and acicular, often acerose, 0.1–0.3 cm × 0.7–1 mm, awns straight, 0.5–1 mm.

greenish to reddish, mostly flat-topped and open to dense;

bracts 3, whorled, short-petiolate, linear-lanceolate to lanceolate, acerose, 0.5–1.5 cm × 3–8(–10) mm, awns straight, 1–3.5 mm.

Involucres

in congested clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4 mm, not corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 0.7–1.5 mm with longer of 3 longest ones more erect than 3 other shorter and less-prominent ones, awns uncinate, 0.5–1 mm.

usually congested in small terminal clusters of 1–3 at node of dichotomies, (4–)5-ribbed, weakly 3-angled, 2–2.5 mm, not corrugate, densely canescent;

teeth (4–)5, essentially erect with longer, prominent, and thickened anterior one 2–4 mm, with straight awn 1–2.5 mm, remaining teeth smaller, 0.5–1 mm, with straight awns 0.3–0.8 mm.

Flowers

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 1/2 their length, dimorphic or sometimes monomorphic, narrowly oblanceolate, apex rounded, those of outer whorl slightly broader and occasionally longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, ovate to oblong, 0.5–0.6 mm.

1, exserted;

perianth, cylindric, 2.5–3.5 mm;

tepals connate 1/2–2/3 their length, dimorphic, entire, those of outer whorl spreading, broadly obovate and rounded apically, those of inner whorl erect, narrowly oblanceolate, 1/2 length of outer ones, acute apically;

stamens slightly exserted;

filaments 2.5–3 mm, glabrous;

anthers yellowish, oblong, 0.5–0.7 mm.

Achenes

brown, lenticular, 3–4 mm.

2.5–3 mm.

2n

= 38.

= (40), 44, (46).

Chorizanthe leptotheca

Chorizanthe spinosa

Phenology Flowering May–Aug. Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, grassland and chaparral communities, pine-oak woodlands Sandy to gravelly flats and slopes, saltbush communities
Elevation (300-)600-1600(-1900) m ((1000-)2000-5200(-6200) ft) 600-1300 m (2000-4300 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe leptotheca is found in the foothills of the San Bernardino Mountains of San Bernardino County southward along the eastern edge of the Santa Ana Mountains, and through the San Jacinto and Santa Rosa mountains of Riverside County into the mountains of central San Diego County. The species is also found in north-central Baja California.

Ramona spineflower is clearly related to Chorizanthe staticoides, but that species occurs to the west of the range of C. leptotheca and the two are not known to be sympatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

G. J. Goodman (1934) referred Chorizanthe spinosa to Eriogonella, but C. spinosa and C. membranacea, the type of Eriogonella, are well isolated from one another, and both are well removed from the remainder of the annual spineflowers. The Mojave spineflower is local and uncommon from southeastern Kern and southern Inyo counties, south into adjacent northeastern Los Angeles and northwestern San Bernardino counties to Antelope and Lucerne valleys.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 463. FNA vol. 5, p. 449.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Quintaria
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms Eriogonella spinosa
Name authority Goodman: Ann. Missouri Bot. Gard. 21: 61. (1934) S. Watson: in W. H. Brewer et al., Bot. California 2: 481. (1880)
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