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Peninsular spineflower, Ramona spineflower

robust spineflower, Scotts Valley spineflower

Habit Plants erect to spreading, 0.5–3(–3.5) × 0.5–3(–5) dm, thinly pubescent. Plants erect to spreading or decumbent, 0.5–3 × 0.1–6 dm, villous.
Leaves

basal;

petiole 1–3(–4) cm;

blade oblong to oblong-ovate, 0.5–2(–3) × 0.3–0.5(–0.7) cm, thinly pubescent adaxially, usually densely tomentose adaxially.

basal or nearly so;

petiole 1–4(–7) cm;

blade oblanceolate, 1.5–5 × 0.2–0.7(–1) cm, villous.

Inflorescences

mostly flat-topped and openly branched, usually reddish;

bracts soon deciduous, 2, occasionally leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, ovate, 0.3–0.4 cm × 2–3 mm, otherwise sessile, linear and acicular, often acerose, 0.1–0.3 cm × 0.7–1 mm, awns straight, 0.5–1 mm.

with secondary branches not suppressed except in terminal clusters of involucres, green to reddish;

bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 1–5 cm × 2–5(–7) mm, awns absent.

Involucres

in congested clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4 mm, not corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 0.7–1.5 mm with longer of 3 longest ones more erect than 3 other shorter and less-prominent ones, awns uncinate, 0.5–1 mm.

1, greenish, cylindric, not ventricose, 2.5–4 mm, with white to pinkish, thin scarious margins restricted to basal portion of teeth, not corrugate, thinly pubescent abaxially;

teeth spreading, equal, 0.3–0.8(–1) mm;

awns uncinate with longer ones 0.7–1.3 mm and anterior one mostly 1–1.3 mm, these alternating with shorter (0.3–0.7 mm) ones.

Flowers

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 1/2 their length, dimorphic or sometimes monomorphic, narrowly oblanceolate, apex rounded, those of outer whorl slightly broader and occasionally longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, ovate to oblong, 0.5–0.6 mm.

slightly exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2.5–4 mm, pubescent abaxially;

tepals connate 1/4 their length, monomorphic, oblanceolate to narrowly oblong, usually truncate to rounded and erose or denticulate apically, occasionally distinctly cuspidate;

stamens 9, included;

filaments distinct, 2–3.5 mm, glabrous;

anthers pink to red or maroon, oblong, 0.6–0.8 mm.

Achenes

brown, lenticular, 3–4 mm.

light brown, globose-lenticular, 3.5–4 mm.

2n

= 38.

Chorizanthe leptotheca

Chorizanthe robusta

Phenology Flowering May–Aug.
Habitat Sandy to gravelly flats and slopes, grassland and chaparral communities, pine-oak woodlands
Elevation (300-)600-1600(-1900) m ((1000-)2000-5200(-6200) ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from USDA
wc Calif
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe leptotheca is found in the foothills of the San Bernardino Mountains of San Bernardino County southward along the eastern edge of the Santa Ana Mountains, and through the San Jacinto and Santa Rosa mountains of Riverside County into the mountains of central San Diego County. The species is also found in north-central Baja California.

Ramona spineflower is clearly related to Chorizanthe staticoides, but that species occurs to the west of the range of C. leptotheca and the two are not known to be sympatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white; plants spreading or decumbent; sandy to gravelly places; Alameda and San Mateo counties s in mountains and near coast to n Monterey County
var. robusta
1. Margins of involucres rose-pink; plants erect; annual grasslands near Scotts Valley, Santa Cruz Mountains, Santa Cruz County
var. hartwegii
Source FNA vol. 5, p. 463. FNA vol. 5, p. 455.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. robusta var. hartwegii, C. robusta var. robusta
Name authority Goodman: Ann. Missouri Bot. Gard. 21: 61. (1934) Parry: Proc. Davenport Acad. Nat. Sci. 5: 176. (1889)
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