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Peninsular spineflower, Ramona spineflower

Douglas' spineflower, San Benito spineflower

Habit Plants erect to spreading, 0.5–3(–3.5) × 0.5–3(–5) dm, thinly pubescent. Plants erect, 1–4(–5) × 0.5–3 dm, villous.
Leaves

basal;

petiole 1–3(–4) cm;

blade oblong to oblong-ovate, 0.5–2(–3) × 0.3–0.5(–0.7) cm, thinly pubescent adaxially, usually densely tomentose adaxially.

basal;

petiole 1–3(–6) cm;

blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous.

Inflorescences

mostly flat-topped and openly branched, usually reddish;

bracts soon deciduous, 2, occasionally leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, ovate, 0.3–0.4 cm × 2–3 mm, otherwise sessile, linear and acicular, often acerose, 0.1–0.3 cm × 0.7–1 mm, awns straight, 0.5–1 mm.

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent.

Involucres

in congested clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4 mm, not corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 0.7–1.5 mm with longer of 3 longest ones more erect than 3 other shorter and less-prominent ones, awns uncinate, 0.5–1 mm.

congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm.

Flowers

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 1/2 their length, dimorphic or sometimes monomorphic, narrowly oblanceolate, apex rounded, those of outer whorl slightly broader and occasionally longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, ovate to oblong, 0.5–0.6 mm.

exserted;

perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire;

stamens 9, slightly exserted;

filaments distinct, 3–4 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

Achenes

brown, lenticular, 3–4 mm.

light brown, globose-lenticular, 3.5–4 mm.

2n

= 38.

= 40.

Chorizanthe leptotheca

Chorizanthe douglasii

Phenology Flowering May–Aug. Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, grassland and chaparral communities, pine-oak woodlands Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands
Elevation (300-)600-1600(-1900) m ((1000-)2000-5200(-6200) ft) (200-)300-1600 m ((700-)1000-5200 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe leptotheca is found in the foothills of the San Bernardino Mountains of San Bernardino County southward along the eastern edge of the Santa Ana Mountains, and through the San Jacinto and Santa Rosa mountains of Riverside County into the mountains of central San Diego County. The species is also found in north-central Baja California.

Ramona spineflower is clearly related to Chorizanthe staticoides, but that species occurs to the west of the range of C. leptotheca and the two are not known to be sympatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 463. FNA vol. 5, p. 452.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms C. nortonii
Name authority Goodman: Ann. Missouri Bot. Gard. 21: 61. (1934) Bentham: Trans. Linn. Soc. London 17: 418. (1836)
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