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fringe spineflower

Turkish rugging

Habit Plants erect to spreading, 1–3(–3.5) × 1–2.5 dm, pubescent and minutely glandular. Plants erect to spreading or ascending, 0.5–6 × 0.5–3(–5) dm, thinly pubescent.
Leaves

basal;

petiole 0.5–3(–5) cm;

blade elliptic to obovate or spatulate, 1–3(–3.5) × 0.2–1(–2.5) cm, thinly pubescent adaxially, sparsely tomentose abaxially.

essentially basal;

petiole 1–3(–4) cm;

blade oblong to oblong-ovate, 0.5–3(–8) × 0.3–1(–2.5) cm, thinly pubescent or glabrous adaxially, usually densely tomentose abaxially.

Inflorescences

open, reddish;

bracts 3 at proximal node, otherwise 2, sessile, scalelike, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 1–2 mm.

mostly flat-topped and open to densely branched, green to gray or reddish;

bracts soon deciduous, 2, occasionally leaflike at proximal nodes and similar to proximal leaf blades only short-petiolate, more reduced and usually sessile, obovate, others linear and acicular, often acerose, (0.1–)0.2–0.5(–1) cm × 1–3(–6) mm, awns straight, 0.5–2 mm.

Involucres

1(–5+), reddish or greenish, cylindric, not ventricose, 4–6(–7) mm, finely corrugate, with thin hyaline margins between teeth, sparsely to densely pubescent;

teeth divergent, unequal, 3 longer ones 1–3 mm, alternating with 3 shorter ones 0.3–1 mm;

awns straight, unequal, 3 longer ones 1–2.5(–3) mm, shorter one (0.3–)0.5–1.5 mm.

usually congested terminally with 1 at node of dichotomies, reddish to purplish, cylindric, not ventricose, 3–4(–5) mm, often irregularly corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 0.7–1.3(–1.5) mm with longest of 3 longer ones more erect than others, these alternating with 3 shorter and less-prominent ones;

awns uncinate, 0.5–1 mm.

Flowers

exserted;

perianth bicolored, with floral tube yellow to yellowish white and tepals white to rose, becoming dark rose to red with age, cylindric, 6–9(–10) mm, glabrous abaxially except for few to several scattered hairs ca. midlength along midribs;

tepals connate 1/2 their length, monomorphic, oblong, fimbriate to laciniate apically;

stamens 9, included;

filaments distinct, 3–8 mm, glabrous;

anthers pink to red, oblong, 0.5–0.7 mm.

mostly included;

perianth rose to red, cylindric, 3–4(–5) mm, pubescent;

tepals connate 1/2 their length, monomorphic to slightly dimorphic, oblong to obovate, rounded to obtuse or truncate apically, occasionally irregularly denticulate, those of outer whorl usually slightly broader and longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 2.5–4(–4.5) mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

Achenes

brown, lenticular, 3–4 mm.

brown, lenticular, 3–4 mm.

2n

= 38, (40, 42).

Chorizanthe fimbriata

Chorizanthe staticoides

Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly or rocky places, coastal scrub, mixed grassland and chaparral communities, pine-oak woodlands
Elevation 300-1700(-1900) m (1000-5600(-6200) ft)
Distribution
from FNA
CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Chorizanthe fimbriata is our only representative of subsect. Flava, a taxon of six species otherwise confined to Baja California. These are the most elegant of the annual species in their remarkable flowers. The flower color, the fine divisions of the tips of the tepals, plus the handsome habit make them of potential horticultural interest for the “need-to-be-challenged” gardeners. In addition to C. fimbriata var. laciniata, C. pulchella Brandegee would be a worthy introduction. Chorizanthe flava Brandegee has bright yellow flowers that contrast dramatically with its reddish mature inflorescence branches. The large (7–9 mm) flowers of C. mutabilis Brandegee are a wonder to behold, the yellow floral tube contrasting with the white to red of the tepals themselves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe staticoides is found in the foothills and mountains of the Coast Ranges from Monterey County southward into San Bernardino, Riverside, and Orange counties. A more insular series of populations occurs on Santa Catalina Island and along the coast and immediately adjacent foothills in Orange and San Diego counties.

Turkish rugging is a highly variable taxon. G. J. Goodman (1934) attempted to parse some of the variants but J. L. Reveal and C. B. Hardham (1989b) merged all of his segregates under a single name. Some of the variation is noteworthy. The insular phase, C. discolor (including C. chrysacantha), is a relatively rare expression restricted to the immediate (C. chrysacantha) and near coastal (C. discolor) mesas and bluffs. Such plants tend to have somewhat longer awns than the inland expression (C. staticoides). Also, the C. chrysacantha phase can be extremely depauperate, resulting in a compact mass overhanging the Pacific Ocean (var. compacta). While the extremes can be noted, useful taxonomic distinctions cannot be made because of intergradation in all features.

E. W. Voegelin (1938) noted the use of an infusion of this plant to treat pimples.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tepals fimbriate apically, distal segments linear to lanceolate, broader than lateral segments; perianths 6-7(-8) mm
var. fimbriata
1. Tepals laciniate apically, distal segments linear, scarcely broader than lateral segments; perianths 8-9(-10) mm
var. laciniata
Source FNA vol. 5, p. 465. FNA vol. 5, p. 462.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. fimbriata var. fimbriata, C. fimbriata var. laciniata
Synonyms C. chrysacantha, C. chrysacantha var. compacta, C. discolor, C. nudicaulis, C. staticoides var. brevispina, C. staticoides subsp. chrysacantha, C. staticoides var. elata, C. staticoides var. latiloba, C. staticoides var. nudicaulis
Name authority Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 17. (1848) Bentham: Trans. Linn. Soc. London 17: 418. (1836)
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