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fringe spineflower

Palmer's spineflower

Habit Plants erect to spreading, 1–3(–3.5) × 1–2.5 dm, pubescent and minutely glandular. Plants erect to spreading, (0.5–)1–3(–4) × 1–3 dm, appressed-pubescent.
Leaves

basal;

petiole 0.5–3(–5) cm;

blade elliptic to obovate or spatulate, 1–3(–3.5) × 0.2–1(–2.5) cm, thinly pubescent adaxially, sparsely tomentose abaxially.

basal;

petiole 1–3 cm;

blade oblanceolate, 1–3 × 0.4–0.8 cm, thinly pubescent.

Inflorescences

open, reddish;

bracts 3 at proximal node, otherwise 2, sessile, scalelike, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 1–2 mm.

with involucres in dense clusters 2–4 cm diam., greenish or reddish to purple;

bracts 2–3 at proximal node, usually leaflike, often with whorl of sessile bracts about midstem, elliptic, 0.5–1.5 cm × 2–6 mm, becoming gradually lanceolate to elliptic, 0.2–1 cm × 1.5–5 mm, at distal nodes scalelike, linear and aciculate, acerose, awns straight, 1–3 mm.

Involucres

1(–5+), reddish or greenish, cylindric, not ventricose, 4–6(–7) mm, finely corrugate, with thin hyaline margins between teeth, sparsely to densely pubescent;

teeth divergent, unequal, 3 longer ones 1–3 mm, alternating with 3 shorter ones 0.3–1 mm;

awns straight, unequal, 3 longer ones 1–2.5(–3) mm, shorter one (0.3–)0.5–1.5 mm.

3–10 or more, reddish to purplish, urceolate, slightly ventricose basally, 3.5–4 mm, without scarious or membranous margins, slightly corrugate, thinly pubescent with slender, curly hairs;

teeth erect to spreading, unequal, 1–2 mm;

awns uncinate, 0.5–1 mm with longer anterior one mostly 1 mm.

Flowers

exserted;

perianth bicolored, with floral tube yellow to yellowish white and tepals white to rose, becoming dark rose to red with age, cylindric, 6–9(–10) mm, glabrous abaxially except for few to several scattered hairs ca. midlength along midribs;

tepals connate 1/2 their length, monomorphic, oblong, fimbriate to laciniate apically;

stamens 9, included;

filaments distinct, 3–8 mm, glabrous;

anthers pink to red, oblong, 0.5–0.7 mm.

exserted;

perianth bicolored with floral tube white to yellow and tepals red, maroon, or dark purple, cylindric, 4–5 mm, glabrous or with few scattered hairs along midrib ca. midlength;

tepals erect, connate 1/2 their length, dimorphic, obovate, those of outer whorl slightly longer than inner whorl, entire, rounded apically, those of inner whorl fimbriate and truncate or somewhat 2-lobed;

stamens 9, exserted;

filaments distinct, 4–5 mm, glabrous;

anthers pink to red or maroon, oblong, 0.9–1 mm.

Achenes

brown, lenticular, 3–4 mm.

brown, globose-lenticular, 3–3.5 mm.

2n

= 38, 40, (48).

Chorizanthe fimbriata

Chorizanthe palmeri

Phenology Flowering May–Aug.
Habitat Gravelly to rocky serpentine and serpentinized igneous outcrops, mixed grassland communities, pine-oak woodlands
Elevation 60-700 m (200-2300 ft)
Distribution
from FNA
CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Chorizanthe fimbriata is our only representative of subsect. Flava, a taxon of six species otherwise confined to Baja California. These are the most elegant of the annual species in their remarkable flowers. The flower color, the fine divisions of the tips of the tepals, plus the handsome habit make them of potential horticultural interest for the “need-to-be-challenged” gardeners. In addition to C. fimbriata var. laciniata, C. pulchella Brandegee would be a worthy introduction. Chorizanthe flava Brandegee has bright yellow flowers that contrast dramatically with its reddish mature inflorescence branches. The large (7–9 mm) flowers of C. mutabilis Brandegee are a wonder to behold, the yellow floral tube contrasting with the white to red of the tepals themselves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe palmeri is known only from the Santa Lucia Mountains, the San Luis Range, and the Huasna area. Populations differ slightly both morphologically and ecologically, but recognition of variants is not suggested. At full anthesis, the reddish stems, involucres, and tepals, plus the localized concentrations of individuals provide for splashes of purplish red on the otherwise grass-brown slopes. Set against the often blackish green of serpentine barrens, the plants can be spotted even from a high-flying aircraft! This species would make an excellent addition to the garden border.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tepals fimbriate apically, distal segments linear to lanceolate, broader than lateral segments; perianths 6-7(-8) mm
var. fimbriata
1. Tepals laciniate apically, distal segments linear, scarcely broader than lateral segments; perianths 8-9(-10) mm
var. laciniata
Source FNA vol. 5, p. 465. FNA vol. 5, p. 457.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. fimbriata var. fimbriata, C. fimbriata var. laciniata
Name authority Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 17. (1848) S. Watson: Proc. Amer. Acad. Arts 12: 271. (1877)
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