Chorizanthe fimbriata |
Chorizanthe howellii |
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fringe spineflower |
Howell's spineflower, Mendocino spineflower |
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Habit | Plants erect to spreading, 1–3(–3.5) × 1–2.5 dm, pubescent and minutely glandular. | Plants spreading or decumbent to somewhat erect, 0.3–1 × 1–5 dm, villous. | ||||
Leaves | basal; petiole 0.5–3(–5) cm; blade elliptic to obovate or spatulate, 1–3(–3.5) × 0.2–1(–2.5) cm, thinly pubescent adaxially, sparsely tomentose abaxially. |
basal or nearly so; petiole 1–4 cm; blade spatulate to broadly obovate, 1–3 × 0.5–1.5(–1.8) cm, villous. |
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Inflorescences | open, reddish; bracts 3 at proximal node, otherwise 2, sessile, scalelike, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 1–2 mm. |
with secondary branches suppressed, greenish to grayish; bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 1–5 cm × 5–15 mm, awns absent. |
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Involucres | 1(–5+), reddish or greenish, cylindric, not ventricose, 4–6(–7) mm, finely corrugate, with thin hyaline margins between teeth, sparsely to densely pubescent; teeth divergent, unequal, 3 longer ones 1–3 mm, alternating with 3 shorter ones 0.3–1 mm; awns straight, unequal, 3 longer ones 1–2.5(–3) mm, shorter one (0.3–)0.5–1.5 mm. |
1, greenish to grayish, broadly cylindric, not ventricose, 3–4 mm, with conspicuous, white, scarious margins between teeth and extending up awn, not corrugate, pubescent; teeth spreading, equal, 0.5–1 mm, awns straight with longer ones 1–2 mm and anterior one mostly 2 mm, these alternating with shorter (0.5–1 mm) ones. |
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Flowers | exserted; perianth bicolored, with floral tube yellow to yellowish white and tepals white to rose, becoming dark rose to red with age, cylindric, 6–9(–10) mm, glabrous abaxially except for few to several scattered hairs ca. midlength along midribs; tepals connate 1/2 their length, monomorphic, oblong, fimbriate to laciniate apically; stamens 9, included; filaments distinct, 3–8 mm, glabrous; anthers pink to red, oblong, 0.5–0.7 mm. |
exserted; perianth bicolored with floral tube white and tepals white to rose, cylindric, (3–)3.5–4.5 mm, pubescent nearly throughout; tepals connate 1/4 their length, dimorphic, oblong, truncate and erose to denticulate apically, those of outer lobes longer and wider than inner ones; stamens 9, included; filaments distinct, 3–4 mm, glabrous; anthers pink to red, oblong, 0.6–0.8 mm. |
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Achenes | brown, lenticular, 3–4 mm. |
light brown, globose-lenticular, 3–4.5 mm. |
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2n | = (72, 74, 76, 78), 80, (82, 84, 86, 88, 90). |
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Chorizanthe fimbriata |
Chorizanthe howellii |
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Phenology | Flowering May–Jul. | |||||
Habitat | Sandy places in coastal dunes and grassland communities | |||||
Elevation | 0-20 m (0-100 ft) | |||||
Distribution |
CA; nw Mexico
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CA |
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Discussion | Varieties 2 (2 in the flora). Chorizanthe fimbriata is our only representative of subsect. Flava, a taxon of six species otherwise confined to Baja California. These are the most elegant of the annual species in their remarkable flowers. The flower color, the fine divisions of the tips of the tepals, plus the handsome habit make them of potential horticultural interest for the “need-to-be-challenged” gardeners. In addition to C. fimbriata var. laciniata, C. pulchella Brandegee would be a worthy introduction. Chorizanthe flava Brandegee has bright yellow flowers that contrast dramatically with its reddish mature inflorescence branches. The large (7–9 mm) flowers of C. mutabilis Brandegee are a wonder to behold, the yellow floral tube contrasting with the white to red of the tepals themselves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Chorizanthe howellii is an octoploid probably derived from an ancient hybrid event involving C. cuspidata var. villosa and perhaps C. valida. It is known only from a dune area north of Fort Bragg in Mendocino County, and is federally listed as endangered. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 465. | FNA vol. 5, p. 456. | ||||
Parent taxa | ||||||
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Name authority | Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 17. (1848) | Goodman: Ann. Missouri Bot. Gard. 21: 44, plate 3, fig. 1. (1934) | ||||
Web links |