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Douglas' spineflower, San Benito spineflower

Santa Barbara spineflower, Wheeler's spineflower

Habit Plants erect, 1–4(–5) × 0.5–3 dm, villous. Plants erect to spreading, 0.5–2(–2.5) × 1–2 dm, thinly pubescent.
Leaves

basal;

petiole 1–3(–6) cm;

blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous.

basal;

petiole 0.5–3 cm;

blade elliptic to oblong, 0.5–2 × 0.2–0.6 cm, thinly pubescent adaxially, tomentose abaxially.

Inflorescences

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent.

mostly flat-topped, openly branched, greenish to reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to leaf blades, short-petiolate, oblong, 0.5–1.2 cm × 2–4 mm, sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.5 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

Involucres

congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm.

in dense terminal clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 2–2.5 mm, corrugate, without scarious or membranous margins, thinly pubescent with stoutish, recurved hairs;

teeth spreading, unequal, 0.3–0.8(–1) mm, with 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.3–0.5 mm.

Flowers

exserted;

perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire;

stamens 9, slightly exserted;

filaments distinct, 3–4 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

exserted;

perianth white or rose or red with white lobes, cylindric, 2.5–3 mm, glabrous except for few scattered hairs ca. midlength along midrib abaxially;

tepals connate 1/2 their length, monomorphic to slightly dimorphic, oblong, rounded apically, those of outer whorl usually slightly broader and longer than those of inner whorl;

stamens 6, included;

filaments distinct, 1.5–2 mm, glabrous;

anthers pink to red, oblong, 0.3–0.4 mm.

Achenes

light brown, globose-lenticular, 3.5–4 mm.

brown, lenticular, 2.5–3 mm.

2n

= 40.

Chorizanthe douglasii

Chorizanthe wheeleri

Phenology Flowering Apr–Jul. Flowering Apr–Jun.
Habitat Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands Gravelly to rocky slopes, coastal scrub communities
Elevation (200-)300-1600 m ((700-)1000-5200 ft) 0-400(-600) m (0-1300(-2000) ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe wheeleri is a rare insular endemic known only from Santa Cruz and Santa Rosa islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 452. FNA vol. 5, p. 464.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. xanti
Synonyms C. nortonii C. insularis
Name authority Bentham: Trans. Linn. Soc. London 17: 418. (1836) S. Watson: Proc. Amer. Acad. Arts 12: 272. (1877)
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