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Douglas' spineflower, San Benito spineflower

one-awn spineflower

Habit Plants erect, 1–4(–5) × 0.5–3 dm, villous. Plants spreading or ascending, 0.2–0.6(–0.8) × 0.5–4(–5) dm, appressed-pubescent.
Leaves

basal;

petiole 1–3(–6) cm;

blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous.

basal;

petiole 0.5–2 cm;

blade oblanceolate, 0.5–1.5(–2) × 0.2–0.8 cm, thinly pubescent.

Inflorescences

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent.

with involucres in small open clusters 0.5–1.5 cm diam., greenish to grayish or reddish;

bracts 2, sessile, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–5 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.4–1.2 cm × 1–2(–3) mm, awns straight, 1.5–4 mm.

Involucres

congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm.

3–10, grayish to reddish, urceolate, slightly ventricose basally, 2–3 mm, without scarious or membranous margins, slightly corrugate, densely grayish-pubescent;

teeth widely spreading to divergent, unequal, 0.3–0.5 or 3–6 mm;

awns straight or uncinate, unequal, with longer anterior one straight, 2.5–5.5 mm, others spreading, uncinate, 0.3–0.5 mm.

Flowers

exserted;

perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire;

stamens 9, slightly exserted;

filaments distinct, 3–4 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

included or only slightly exserted;

perianth bicolored with floral tube greenish white and tepals white, cylindric, 2–3 mm, sparsely pubescent;

tepals connate 2/3 their length, dimorphic, linear-oblong, those of outer whorl spreading, narrowly oblong, 1.5 times longer than those of inner whorl, rounded but with minute cusp or 3 teeth apically, those of inner whorl erect to slightly spreading, acute, entire apically;

stamens 3, included;

filaments distinct, 1–2 mm, glabrous;

anthers white, ovate, 0.4–0.5 mm.

Achenes

light brown, globose-lenticular, 3.5–4 mm.

brown, globose-lenticular, 2–3 mm.

2n

= 40.

= (78), 80, (82).

Chorizanthe douglasii

Chorizanthe uniaristata

Phenology Flowering Apr–Jul. Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands Sandy to gravelly talus or clay flats and slopes, mixed grassland and chaparral communities, pine-oak woodlands
Elevation (200-)300-1600 m ((700-)1000-5200 ft) 800-1900 m (2600-6200 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe uniaristata is scattered in the Inner Coast Ranges and across the Transverse and Tehachapi ranges to the southern Sierra Nevada.

One-awn spineflower is a polyploid, but whether an autopolyploid or an autoallopolyploid has not been determined. It has the smallest meiotic chromosomes observed by C. B. Hardham (1989).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 452. FNA vol. 5, p. 460.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms C. nortonii
Name authority Bentham: Trans. Linn. Soc. London 17: 418. (1836) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 195. (1870)
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