Chorizanthe douglasii |
Chorizanthe rigida |
|
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Douglas' spineflower, San Benito spineflower |
devil's spineflower, rigid spineflower, rigid spiny-herb, spiny-herb |
|
Habit | Plants erect, 1–4(–5) × 0.5–3 dm, villous. | Plants erect, 0.2–0.8(–1.5) × 0.1–0.7(–1) dm, pubescent. |
Leaves | basal; petiole 1–3(–6) cm; blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous. |
basal and cauline; petiole 0.5–3(–4) cm; basal blade broadly elliptic to obovate, 0.5–2.5 × (0.3–)0.5–2 cm, thinly pubescent adaxially, more densely so to tomentose abaxially; proximal cauline leaf soon deciduous, 1, blade similar to basal leaf blades only 1–2(–2.5) × 0.5–1.5 cm, mucronate to awn-tipped, awn mostly 2–4 mm; distal cauline leaf blade persistent, 1 per node, sessile, blade linear to linear-lanceolate, 0.1–1.5 × 0.05–0.15 cm, becoming hard and thornlike with age. |
Inflorescences | cymose, dichotomously branched throughout, white to greenish or reddish; bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent. |
with involucres in dense clusters in axils of bracts, these on short shoots and each subtended by cauline leaves; bracts 2, subopposite to opposite, linear, 0.5–1(–1.2) cm × 1–2 mm, awns straight, 2–4 mm. |
Involucres | congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent; teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm. |
1, greenish, urceolate, 3-ribbed, 2–3 mm, corrugate, pubescent, rarely villous near base in some; teeth 3, unequal, with thickened anterior tooth toward base, 5–10 mm, sometimes expanding and becoming lanceolate to narrowly elliptic, others 0.5–1.2 mm; awns straight. |
Flowers | exserted; perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially; tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire; stamens 9, slightly exserted; filaments distinct, 3–4 mm, glabrous; anthers pink to red, oblong, 0.5–0.6 mm. |
1–2, included to slightly exserted; perianth yellow, cylindric, 1.5–1.8 mm, densely pubescent abaxially; tepals connate ca. 2/3 their length, monomorphic, oblong, rounded, entire apically; stamens 9, slightly exserted; filaments distinct, 0.5–1 mm, glabrous; anthers yellowish, ovate, 0.2–0.3 mm. |
Achenes | light brown, globose-lenticular, 3.5–4 mm. |
brown, 3-gonous, (1.5–)1.8–2.2 mm. |
2n | = 40. |
= 38, 40. |
Chorizanthe douglasii |
Chorizanthe rigida |
|
Phenology | Flowering Apr–Jul. | Flowering Feb–Jun. |
Habitat | Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands | Sandy to gravelly or rocky flats and slopes, desert scrub |
Elevation | (200-)300-1600 m ((700-)1000-5200 ft) | -60-1900 m. (-200-6200 ft.) |
Distribution |
CA
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AZ; CA; NV; UT; Mexico (Baja California, Sonora)
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Discussion | Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Anyone with the misfortune to step bare-footed on Chorizanthe rigida after the plant has dried instantly appreciates its common name. The species is widespread on the Mojave and Sonoran deserts but only occasionally is it locally abundant or weedy. It is found also along the Lahontan Trough in western Nevada, a well-known biogeographic extension route north of the Mojave Desert (J. L. Reveal 1980). The exceedingly compact and dense inflorescences with suppressed secondary branches result in a series of leaves and bracts that subtend a closely arranged series of bracteated and involucrated flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 452. | FNA vol. 5, p. 467. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. nortonii | Acanthogonum rigidum |
Name authority | Bentham: Trans. Linn. Soc. London 17: 418. (1836) | (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870) |
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