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Douglas' spineflower, San Benito spineflower

prickly spineflower, straight-awn spineflower

Habit Plants erect, 1–4(–5) × 0.5–3 dm, villous. Plants spreading to decumbent, 0.3–0.8(–1) × 0.5–4(–5) dm, appressed-pubescent.
Leaves

basal;

petiole 1–3(–6) cm;

blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous.

basal;

petiole 0.5–2 cm;

blade oblanceolate to spatulate, 0.5–1.5(–2) × 0.2–0.6 cm, thinly pubescent.

Inflorescences

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent.

with involucres in small, open clusters 0.5–1.5 cm diam., greenish to grayish;

bracts 2, without whorl of sessile bracts about midstem, usually leaflike, oblanceolate to elliptic, 0.5–1.5 cm × 1.5–5 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.3–0.8 cm × 1–2 mm, awns straight, 0.5–1.5 mm.

Involucres

congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm.

3–10+, grayish to reddish, urceolate, slightly ventricose basally, 2–2.5(–3) mm, slightly corrugate, without scarious or membranous margins, densely pubescent;

teeth spreading, unequal, 1–2 mm;

awns straight or uncinate, unequal, with longer anterior one straight, mostly 1.5–2.5 mm, others uncinate, 0.3–0.6 mm.

Flowers

exserted;

perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire;

stamens 9, slightly exserted;

filaments distinct, 3–4 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

exserted;

perianth bicolored with floral tube yellow and tepals yellow or white, cylindric, 3.5–4 mm, sparsely pubescent;

tepals connate 1/2 their length, dimorphic, obovate, those of outer whorl white, obovate to nearly orbiculate, 3–4 times longer than those of inner whorl, , truncate to slightly 2-lobed apically, those of inner lobes erect, yellow, broadly obovate, truncate and erose apically;

stamens 9, included;

filaments distinct, 1–1.5 mm, glabrous;

anthers yellow to golden, oblong, 0.5–0.6 mm.

Achenes

light brown, globose-lenticular, 3.5–4 mm.

brown, globose-lenticular, 3–3.5 mm.

2n

= 40.

= (36), 40, (44).

Chorizanthe douglasii

Chorizanthe rectispina

Phenology Flowering Apr–Jul. Flowering May–Jul.
Habitat Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands Sandy to gravelly flats and slopes, mixed grassland communities, pine-oak woodlands
Elevation (200-)300-1600 m ((700-)1000-5200 ft) 200-600 m (700-2000 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chorizanthe rectispina is infrequent and localized in the Coast Ranges of west-central California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 452. FNA vol. 5, p. 460.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Synonyms C. nortonii
Name authority Bentham: Trans. Linn. Soc. London 17: 418. (1836) Goodman: Ann. Missouri Bot. Gard. 21: 72. (1934)
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