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Douglas' spineflower, San Benito spineflower

Monterey spineflower

Habit Plants erect, 1–4(–5) × 0.5–3 dm, villous. Plants prostrate to ascending or erect, 0.5–2(–2.5) × 0.5–10 dm, grayish-villous.
Leaves

basal;

petiole 1–3(–6) cm;

blade oblanceolate, 0.5–2(–4) × 0.1–0.4(–1) cm, villous.

basal;

petiole (0.5–)1–3(–4) cm;

blade oblanceolate, (0.5–)1–5(–7) × (0.3–)0.4–0.7(–1) cm, villous.

Inflorescences

cymose, dichotomously branched throughout, white to greenish or reddish;

bracts usually 2, similar to proximal leaf blades, typically with whorl of 3–5 ca. midstem, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–2(–3) cm × 1–5(–10) mm, awns absent.

rather dense with secondary branches suppressed, grayish;

bracts 2, similar to leaf blades at proximal nodes only reduced, short-petiolate, becoming linear and aciculate at distal nodes, acerose, 0.5–7 cm × 2–7 mm, awns 0.5–1.2 mm.

Involucres

congested in small leafy terminal clusters of 2–4 at nodes of dichotomies, greenish, cylindric, slightly ventricose basally, 3–5 mm, with conspicuous, purple, broad, membranous margins typically extending across sinuses, finely corrugated, hispid at least along ridges, otherwise sparsely pubescent;

teeth spreading, equal, (0.7–)1–1.5 mm, awns straight, 0.5–1 mm.

1, grayish, cylindric, often ventricose basally, 2–2.5(–3) mm, with distinct, white to pink or purple, scarious margins extending nearly full length of awn, corrugate, villous abaxially;

teeth spreading, equal, 0.5–1.5 mm;

awns uncinate with longer ones 2–3 mm and alternating with shorter (1–1.5 mm) ones.

Flowers

exserted;

perianth white to rose, cylindric, 3.5–4(–4.5) mm, slightly pubescent abaxially;

tepals connate 2/3 their length, monomorphic, obovate, 2-lobed or denticulate apically, infrequently inner whorl entire;

stamens 9, slightly exserted;

filaments distinct, 3–4 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3.5 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, obovate to oblong, acute to truncate and erose apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, ovate, 0.3–0.4 mm.

Achenes

light brown, globose-lenticular, 3.5–4 mm.

dark brown, globose-lenticular, 2–2.5 mm.

2n

= 40.

Chorizanthe douglasii

Chorizanthe pungens

Phenology Flowering Apr–Jul.
Habitat Sandy to gravelly flats and slopes, mixed grassland communities, oak and pine woodlands
Elevation (200-)300-1600 m ((700-)1000-5200 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Chorizanthe douglasii is restricted to the Santa Lucia Mountains and to the San Gabilan and La Panza ranges of west-central California. The species is infrequent but can be locally common. A single collection made in the Santa Cruz Mountains (Rowntree s.n., 16 Jun 1929, CAS) may have been made in Santa Cruz County, but the location is uncertain and no other collection is known from that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of involucres white (rarely pinkish), scarious; plants prostrate to slightly ascending; coastal areas and adjacent inland valleys
var. pungens
1. Margins of involucres dark pinkish to purple, scarious; plants slightly ascending to erect; coastal mountains
var. hartwegiana
Source FNA vol. 5, p. 452. FNA vol. 5, p. 453.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. pungens var. hartwegiana, C. pungens var. pungens
Synonyms C. nortonii
Name authority Bentham: Trans. Linn. Soc. London 17: 418. (1836) Bentham: Trans. Linn. Soc. London 17: 419, plate 19, fig. 2. (1836)
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