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San Francisco spineflower

wrinkled spineflower

Habit Plants decumbent to prostrate or ascending, 0.5–2(–2.5) × 0.5–10 dm, villous. Plants erect, 0.3–1.5 × 0.3–1 dm, thinly tomentose.
Leaves

basal;

petiole (0.5–)1–3 cm;

blade oblanceolate, (0.5–)1–5 × (0.3–)0.4–0.7(–1) cm, villous.

basal or nearly so;

petiole 0.5–2(–3) cm;

blade round-ovate, (0.5–)0.8–1.5(–2) × (0.3–)0.5–1.5(–2) cm, thinly floccose to tomentose.

Inflorescences

rather dense with secondary branches suppressed, greenish to reddish;

bracts 2, similar to proximal leaf blades only reduced, short-petiolate, becoming narrowly elliptic to linear-lanceolate and aciculate at distal nodes, acerose, 0.5–5 cm × 2–7 mm, awns 0.5–1.2 mm.

with involucres in small clusters 0.5–1 cm diam., green to tan or reddish;

bracts 2, linear to linear-lanceolate, acicular, 2–7 cm × 1–2.5 mm, awns slightly curved, 0.5–1 mm.

Involucres

1, greenish, cylindric, often ventricose basally, 1–3 mm, without scarious margins or if so then white to pink, thin, and restricted to basal portions of teeth, corrugate, villous abaxially;

teeth spreading, equal, 0.5–2 mm;

awns uncinate or straight with longer ones 2–3 mm and anterior one mostly 2.5–3 mm, these alternating with shorter 1–1.5(–1.7) mm ones.

1, green to tan, cylindric, 3-angled but 3-ribbed, 3–4 mm, markedly transverse corrugate, glabrate;

teeth 3, equal, 2–4.5 mm;

awns uncinate, 0.6–1 mm.

Flowers

included to slightly exserted;

perianth bicolored with floral tube white and tepals white to rose, cylindric, 2–3 mm, pubescent abaxially;

tepals connate less than 1/4 their length, monomorphic, oblong, truncate to 3-lobed and distinctly cuspidate apically;

stamens 9, slightly exserted;

filaments distinct, 2–3 mm, glabrous;

anthers cream to rose, narrowly oblong, 0.3–0.4 mm.

1, included to slightly exserted;

perianth white, cylindric, 2–2.5 mm, thinly pubescent abaxially;

tepals connate ca. 2/3 their length, monomorphic, oblong, acute, entire apically;

stamens 6, slightly exserted;

filaments distinct, 0.8–1 mm, glabrous;

anthers cream, ovate, 0.4–0.5 mm.

Achenes

light brown, globose-lenticular, 2–3 mm.

brown, lenticular, 2.5–3 mm.

2n

= 38.

Chorizanthe cuspidata

Chorizanthe corrugata

Phenology Flowering Feb–May.
Habitat Sandy to gravelly flats and slopes, mixed grassland, saltbush, creosote bush, and sagebrush communities
Elevation -70-1000 m (-200-3300 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Chorizanthe corrugata is found mainly in the Mojave and Sonoran deserts. The narrow, transversely corrugated involucral tube is diagnostic. Some anomalous flowers with four or eight stamens have been seen but this condition was always associated with other flowers bearing the normal number.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral awns uncinate apically; San Francisco area
var. cuspidata
1. Involucral awns straight, rarely curved or with 1 awn uncinate; Marin and Sonoma counties
var. villosa
Source FNA vol. 5, p. 454. FNA vol. 5, p. 468.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Acanthogonum
Sibling taxa
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. cuspidata var. cuspidata, C. cuspidata var. villosa
Synonyms C. pungens var. cuspidata Acanthogonum corrugatum
Name authority S. Watson: Proc. Amer. Acad. Arts 17: 379. (1882) (Torrey) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 198. (1870)
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