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Blakley's spineflower

riverside spineflower, Xantus' spineflower

Habit Plants spreading to ascending, 0.5–1.5 × 0.5–3 dm, thinly pubescent. Plants erect to infrequently spreading, (0.3–)0.5–2.5(–3) × 0.5–3(–5) dm, thinly pubescent.
Leaves

basal;

petiole 0.5–2 cm;

blade oblanceolate, 0.5–2.5 × 0.3–0.8 cm, thinly pubescent.

basal or nearly so;

petiole 1–2(–3) cm;

blade oblong or oblong-ovate to ovate, 0.3–1(–1.5) × 0.3–0.8(–1) cm, thinly pubescent adaxially, densely tomentose abaxially.

Inflorescences

with involucres in dense clusters 1–2 cm diam., yellowish green;

bracts 2, without whorl of sessile bracts about midstem, usually leaflike, oblanceolate, 0.5–1.5 cm × 1.5–3 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.3–0.8 cm × 1–2 mm, awns straight, 1–2.5 mm.

mostly flat-topped and openly branched, usually reddish;

bracts persistent, 2, usually leaflike at proximal nodes and similar to proximal leaf blades only more reduced, short-petiolate, oblong-ovate to ovate, 0.3–0.8 cm × 2–6 mm, becoming sessile, reduced and scalelike at distal nodes, linear, acicular, often acerose, 0.1–0.4 cm × 0.5–1 mm, awns straight, 0.5–1 mm.

Involucres

3–10+, yellowish green, urceolate, slightly ventricose basally, 3–4.5 mm, slightly corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight or slightly curved, mostly 2 mm, others uncinate, 0.5–1.5 mm.

in open clusters with 1 at node of dichotomies, reddish, cylindric, not ventricose, 3–4.5 mm, not corrugate, without scarious or membranous margins, thinly to densely pubescent;

teeth spreading, unequal, 0.7–1.5 mm, 3 longer ones more erect than 3 shorter and less-prominent ones;

awns uncinate, 0.5–1 mm.

Flowers

exserted;

perianth bicolored with floral tube greenish white to white and the lobes white to pinkish, cylindric, 5–6 mm, sparsely pubescent;

tepals connate 2/3 their length, dimorphic, obovate, those of outer whorl erect, slightly longer than those of inner whorl, 2-lobed apically, those of inner whorl erect, 2-lobed, erose apically;

stamens 9, included;

filaments distinct, 5–5.5 mm, glabrous;

anthers yellow to golden, oblong, 1–1.2 mm.

long-exserted;

perianth rose to red, infrequently with white lobes, cylindric, 4.5–6 mm, pubescent;

tepals connate ca. 2/3 their length, monomorphic to slightly dimorphic, narrowly oblanceolate, rounded apically, those of outer whorl occasionally slightly broader and longer than those of inner whorl;

stamens 9, mostly included;

filaments distinct, 4–6 mm, glabrous;

anthers pink to red, oblong, 0.5–0.6 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

brown, lenticular, 4–4.5 mm.

2n

= ca. 38.

Chorizanthe blakleyi

Chorizanthe xanti

Phenology Flowering May–Jul.
Habitat Sandy or gravelly flats and slopes, chaparral communities, oak woodlands
Elevation 600-1600 m (2000-5200 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chorizanthe blakleyi is known only from north-facing slopes and foothills of the Sierra Madre. The species is rare and localized. The yellowish green stem, branches, and involucres readily distinguish it. The white flowers align it with C. obovata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres thinly pubescent
var. xanti
1. Involucres densely white-pubescent
var. leucotheca
Source FNA vol. 5, p. 459. FNA vol. 5, p. 463.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala
Sibling taxa
C. angustifolia, C. biloba, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri
Subordinate taxa
C. xanti var. leucotheca, C. xanti var. xanti
Name authority Hardham: Leafl. W. Bot. 10: 95. (1964) S. Watson: Proc. Amer. Acad. Arts 12: 272. (1877)
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