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Blakley's spineflower

Habit Plants spreading to ascending, 0.5–1.5 × 0.5–3 dm, thinly pubescent. Plants prostrate to spreading or erect, mostly thinly pubescent.
Stems

sometimes disarticulating at each node.

Leaves

basal;

petiole 0.5–2 cm;

blade oblanceolate, 0.5–2.5 × 0.3–0.8 cm, thinly pubescent.

blades linear to lanceolate, obovate, round, or spatulate.

Inflorescences

with involucres in dense clusters 1–2 cm diam., yellowish green;

bracts 2, without whorl of sessile bracts about midstem, usually leaflike, oblanceolate, 0.5–1.5 cm × 1.5–3 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.3–0.8 cm × 1–2 mm, awns straight, 1–2.5 mm.

bracts mostly 2, opposite, scalelike or if leaflike then similar to basal leaves only reduced, occasionally deciduous in early anthesis, with or without awns.

Involucres

3–10+, yellowish green, urceolate, slightly ventricose basally, 3–4.5 mm, slightly corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight or slightly curved, mostly 2 mm, others uncinate, 0.5–1.5 mm.

cylindric to narrowly turbinate, campanulate, or urceolate, occasionally ventricose basally, 3-,5-, or 6-toothed, with or without membranous or scarious margins;

teeth erect to spreading or divergent, connate at least 1/2 their length, typically shallow, mostly unequal, with alternating long and short awns, often with anterior one longest.

Flowers

exserted;

perianth bicolored with floral tube greenish white to white and the lobes white to pinkish, cylindric, 5–6 mm, sparsely pubescent;

tepals connate 2/3 their length, dimorphic, obovate, those of outer whorl erect, slightly longer than those of inner whorl, 2-lobed apically, those of inner whorl erect, 2-lobed, erose apically;

stamens 9, included;

filaments distinct, 5–5.5 mm, glabrous;

anthers yellow to golden, oblong, 1–1.2 mm.

1(–2), white to pink or rose, maroon or purple, or yellow, thinly pubescent at least along midribs abaxially;

stamens 3–9;

filaments adnate at base of floral tube or faucially;

filaments sometimes connate into short tube.

Achenes

brown, globose-lenticular, 3–3.5 mm.

brown, lenticular or globose-lenticular, or 3-gonous.

Seeds

embryo straight or rarely curved.

2n

= ca. 38.

Chorizanthe blakleyi

Chorizanthe subg. Amphietes

Phenology Flowering May–Jul.
Habitat Sandy or gravelly flats and slopes, chaparral communities, oak woodlands
Elevation 600-1600 m (2000-5200 ft)
Distribution
from FNA
CA
[BONAP county map]
w United States; nw Mexico; sw South America
Discussion

Of conservation concern.

Chorizanthe blakleyi is known only from north-facing slopes and foothills of the Sierra Madre. The species is rare and localized. The yellowish green stem, branches, and involucres readily distinguish it. The white flowers align it with C. obovata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 39 (31 in the flora).

Most species of subg. Amphietes are found in California. Of the others, one is known only from southernmost Peru to central Chile (Chorizanthe commissuralis J. Rémy), while the rest are known only from Baja California, Mexico. Those include C. inequalis S. Stokes, C. turbinata Wiggins, C. mutabilis Brandegee, C. rosulenta Reveal, C. pulchella Brandegee, C. flava Brandegee, and C. interposita Goodman. The latter is the only member of sect. Clastoscapa, the only section of subg. Amphietes not found in our flora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 459. FNA vol. 5, p. 450.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe
Sibling taxa
C. angustifolia, C. biloba, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
Name authority Hardham: Leafl. W. Bot. 10: 95. (1964) Reveal & Hardham: Phytologia 66: 113. (1989)
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