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Blakley's spineflower

brittle spine flower

Habit Plants spreading to ascending, 0.5–1.5 × 0.5–3 dm, thinly pubescent. Plants spreading to erect, 0.5–3(–5) × 0.5–3 dm, thinly pubescent, often with appressed hairs, infrequently somewhat strigose or glabrate.
Leaves

basal;

petiole 0.5–2 cm;

blade oblanceolate, 0.5–2.5 × 0.3–0.8 cm, thinly pubescent.

basal;

petiole 0.5–2 cm;

blade oblanceolate to narrowly elliptic or spatulate, (1–)1.5–3(–4) × 0.1–1 cm, pubescent.

Inflorescences

with involucres in dense clusters 1–2 cm diam., yellowish green;

bracts 2, without whorl of sessile bracts about midstem, usually leaflike, oblanceolate, 0.5–1.5 cm × 1.5–3 mm, gradually reduced and becoming scalelike at distal nodes, linear, aciculate, acerose, 0.3–0.8 cm × 1–2 mm, awns straight, 1–2.5 mm.

green;

bracts 2, similar to proximal leaf blades only more reduced, 0.3–1(–1.5) cm × 1–2.5 mm, becoming sessile and scalelike at distal nodes, linear, acicular, awns 0.2–0.5 mm.

Involucres

3–10+, yellowish green, urceolate, slightly ventricose basally, 3–4.5 mm, slightly corrugate, without scarious or membranous margins, thinly pubescent;

teeth spreading, unequal, 1–3 mm;

awns straight or uncinate with longer anterior one straight or slightly curved, mostly 2 mm, others uncinate, 0.5–1.5 mm.

1, green, 3–5 mm, not corrugate, thinly strigose;

teeth divergent, 0.4–1.2 mm;

awns uncinate, 0.2–0.5 mm.

Flowers

exserted;

perianth bicolored with floral tube greenish white to white and the lobes white to pinkish, cylindric, 5–6 mm, sparsely pubescent;

tepals connate 2/3 their length, dimorphic, obovate, those of outer whorl erect, slightly longer than those of inner whorl, 2-lobed apically, those of inner whorl erect, 2-lobed, erose apically;

stamens 9, included;

filaments distinct, 5–5.5 mm, glabrous;

anthers yellow to golden, oblong, 1–1.2 mm.

included;

perianth greenish white to white or pale yellowish white, cylindric, 2–4 mm;

tepals connate 3/4 their length, monomorphic, linear to narrowly oblanceolate, acute, entire apically;

stamens slightly exserted;

filaments distinct, 2–3.5 mm, glabrous;

anthers white to pale yellow, ovate, 0.3–0.4 mm.

Achenes

brown, globose-lenticular, 3–3.5 mm.

dark brown, lenticular, 3–4 mm.

2n

= ca. 38.

Chorizanthe blakleyi

Chorizanthe brevicornu

Phenology Flowering May–Jul.
Habitat Sandy or gravelly flats and slopes, chaparral communities, oak woodlands
Elevation 600-1600 m (2000-5200 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
AZ; CA; ID; NV; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Chorizanthe blakleyi is known only from north-facing slopes and foothills of the Sierra Madre. The species is rare and localized. The yellowish green stem, branches, and involucres readily distinguish it. The white flowers align it with C. obovata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Chorizanthe brevicornu has stems and branches that easily disarticulate at the nodes. Dried specimens often are reduced to a mere jumble without careful handling. The vegetative fragments will not regenerate new plants, but the involucres (each with a single flower bearing a single achene) easily disarticulate from the parent plant, and with the aid of the awns on the teeth of the involucre, may be readily distributed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades oblanceolate to narrowly elliptic, 0.1-0.3(-0.5) cm wide; involucres prominently ribbed at maturity; Mojave and Sonoran deserts
var. brevicornu
1. Leaf blades broadly oblanceolate to broadly spatulate, 0.5-1 cm wide; involucres obscurely ribbed at maturity; Great Basin Desert and Snake River Plains
var. spathulata
Source FNA vol. 5, p. 459. FNA vol. 5, p. 469.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Ptelosepala Polygonaceae > subfam. Eriogonoideae > Chorizanthe > subg. Amphietes > sect. Fragile
Sibling taxa
C. angustifolia, C. biloba, C. brevicornu, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
C. angustifolia, C. biloba, C. blakleyi, C. breweri, C. clevelandii, C. corrugata, C. cuspidata, C. diffusa, C. douglasii, C. fimbriata, C. howellii, C. leptotheca, C. membranacea, C. obovata, C. orcuttiana, C. palmeri, C. parryi, C. polygonoides, C. procumbens, C. pungens, C. rectispina, C. rigida, C. robusta, C. spinosa, C. staticoides, C. stellulata, C. uniaristata, C. valida, C. ventricosa, C. watsonii, C. wheeleri, C. xanti
Subordinate taxa
C. brevicornu var. brevicornu, C. brevicornu var. spathulata
Name authority Hardham: Leafl. W. Bot. 10: 95. (1964) Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 177. (1859)
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