Cheilanthes wootonii |
Cheilanthes intertexta |
|
---|---|---|
beaded lipfern, Wooton's lip fern |
coastal lip fern |
|
Stems | long-creeping, 1–3 mm diam.; scales uniformly brown or weakly bicolored with poorly defined, dark, central stripe, lanceolate-ovate, straight to slightly contorted, loosely appressed, often deciduous on older portions of stem. |
short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
Leaves | scattered, 7–35 cm; vernation noncircinate. |
clustered, 4–25 cm; vernation noncircinate. |
Petiole | usually dark brown, rounded adaxially. |
dark brown, rounded adaxially. |
Blade | oblong-lanceolate, 3–4-pinnate at base, 2–5 cm wide; rachis rounded adaxially, with scattered linear-lanceolate scales and sparse monomorphic pubescence. |
lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. |
Ultimate segments | round to oblong, beadlike, the largest 1–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous. |
oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
± continuous around segment margins. |
Sporangia | containing 32 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, lanceolate-ovate, truncate or subcordate at base, without overlapping basal lobes, conspicuous, the largest 0.4–0.8 mm wide, strongly imbricate, often concealing ultimate segments, ciliate, with coarse cilia often confined to proximal 1/2. |
green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. |
n | = 2n = 90, apogamous. |
|
Cheilanthes wootonii |
Cheilanthes intertexta |
|
Phenology | Sporulating summer–fall. | Sporulating late spring–fall. |
Habitat | Rocky slopes and ledges, usually on igneous substrates | Rocky slopes and ledges, usually on igneous substrates |
Elevation | 800–2900 m (2600–9500 ft) | 500–2800 m (1600–9200 ft) |
Distribution |
AZ; CA; CO; NM; NV; OK; TX; UT; n Mexico
|
CA; NV; OR
|
Discussion | Like its close relative Cheilanthes lindheimeri, C. wootonii is an apogamous triploid of unknown parentage. With the recognition of C. yavapensis as a distinct species, the name C. wootonii is restricted to populations with leaf blades that appear glabrous adaxially, costal scales that are often ciliate only in the proximal half, and stem scales that are usually brown and loosely appressed. In addition, C. wootonii is distinguished from C. yavapensis by having smaller spores, averaging less than 62 µm in diameter. These characteristics can be subtle, and some specimens will be difficult to place in either C. wootonii or C. yavapensis. T. Reeves (1979) identified several specimens from Arizona that he hypothesized were hybrids between C. wootonii and C. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | C. covillei subsp. intertexta | |
Name authority | Maxon: Proc. Biol. Soc. Wash. 3: 146. (1918) | (Maxon) Maxon: in Abrams, Ill. Fl. Pacific States 1: 28. (1923) |
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