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viscid lip fern

lace fern, lace lipfern

Stems

short-creeping, usually 4–8 mm diam.;

scales uniformly brown, linear-subulate, strongly contorted, loosely appressed, persistent.

short-creeping, 4–8 mm diam.;

scales uniformly brown or with poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent.

Leaves

clustered, 6–30 cm;

vernation circinate.

clustered, 5–25 cm;

vernation noncircinate.

Petiole

dark brown, flattened or slightly grooved distally on adaxial surface.

dark brown, rounded adaxially.

Blade

narrowly oblong to linear, 3-pinnate-pinnatifid at base, 1–4 cm wide;

rachis flattened or slightly grooved adaxially, lacking scales, with monomorphic pubescence.

linear-oblong, 2–3-pinnate at base, 1–2.5 cm wide;

rachis rounded adaxially, with scattered linear scales, not pubescent.

Ultimate segments

oblong to lanceolate, not beadlike, the largest 3–4 mm, abaxially and adaxially glandular-pubescent with short, sticky, capitate glands.

oblong or rarely oval, beadlike, the largest 1.5–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered, branched hairs or glabrescent.

Pinnae

not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing glandular pubescent adaxially.

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing sparsely pubescent or glabrous adaxially.

False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.

marginal, slightly differentiated, 0.05–0.25 mm wide.

Sori

usually discontinuous, concentrated on apical and lateral lobes.

± continuous around segment margins.

Sporangia

containing 64 spores.

containing 64 spores.

Costae

green adaxially for most of length;

abaxial scales absent.

green adaxially for most of length;

abaxial scales multiseriate, linear, truncate at base, inconspicuous, the largest 0.1–0.4 mm wide, loosely imbricate, not concealing ultimate segments, long-ciliate, cilia usually confined to base.

Cheilanthes viscida

Cheilanthes gracillima

Phenology Sporulating late spring–fall. Sporulating summer–fall.
Habitat Cliffs and rocky slopes, usually on igneous substrates Cliffs and rocky slopes, usually on igneous substrates
Elevation 200–1300 m (700–4300 ft) 800–3000 m (2600–9800 ft)
Distribution
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
from FNA
CA; ID; MT; NV; OR; UT; WA; AB; BC
[WildflowerSearch map]
Discussion

Cheilanthes viscida is confined to a relatively small region in the deserts of California. Variations in spore size among populations suggest that the species may include more than one cytotype.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cheilanthes gracillima is a well-marked species, but it apparently hybridizes with C. intertexta (see reticulogram) to produce plants of intermediate morphology with malformed spores that have been called C. gracillima var. aberrans M. E. Jones (A. R. Smith 1974).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. wootonii, C. wrightii, C. yavapensis
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. covillei, C. eatonii, C. feei, C. fendleri, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
Synonyms Myriopteris gracillima
Name authority Davenport: Bull. Torrey Bot. Club 6: 191. (1877) D. C. Eaton: in Emory, Rep. U.S. Mex. Bound. 2: 234. (1859)
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