Cheilanthes lanosa |
Cheilanthes yavapensis |
|
---|---|---|
hairy lip fern |
graceful lipfern, Yavapai lip fern |
|
Stems | compact to short-creeping, usually 4–8 mm diam.; scales often uniformly brown but at least some on each plant with thin, poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
long-creeping, 1–3 mm diam.; scales often bicolored, with broad, poorly defined, dark, central stripe and narrow, brown margins, lanceolate, straight to slightly contorted, strongly appressed, persistent. |
Leaves | clustered, 7–50 cm; vernation circinate. |
scattered, 7–35 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
dark brown, rounded adaxially. |
Blade | linear-oblong to lanceolate, usually 2-pinnate-pinnatifid at base, 1.5–5 cm wide; rachis rounded adaxially, lacking scales, with monomorphic pubescence. |
oblong-lanceolate to nearly ovate, 4-pinnate at base, 2–6 cm wide; rachis rounded adaxially, with scattered linear-lanceolate scales and sparse monomorphic pubescence. |
Ultimate segments | oblong to lanceolate, not beadlike, the largest 3–5 mm, abaxially and adaxially sparsely hirsute with long, segmented hairs. |
round to oblong, beadlike, the largest usually 1–2 mm, abaxially glabrous or with a few small scales near base, adaxially appearing sparsely pubescent but actually nearly glabrous. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing sparsely hirsute adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing sparsely pubescent adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | discontinuous, concentrated on small apical and lateral lobes. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 32 spores. |
Costae | brown adaxially for most of length; abaxial scales absent. |
green adaxially for most of length; abaxial scales multiseriate, lanceolate, truncate to cordate at base, without overlapping basal lobes, conspicuous, the largest 0.4–1 mm wide, strongly imbricate, often concealing ultimate segments, ciliate, cilia coarse, usually distributed entire length of scale. |
n | = 2n = 120, apogamous. |
|
2n | = 60. |
|
Cheilanthes lanosa |
Cheilanthes yavapensis |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, on a variety of substrates including limestone and granite | Rocky slopes and ledges, usually on igneous substrates |
Elevation | 100–800 m (300–2600 ft) | 500–2400 m (1600–7900 ft) |
Distribution |
AL; AR; CT; FL; GA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV |
AZ; NM; TX |
Discussion | Cheilanthes lanosa is apparently confined to the forests and prairies of eastern North America, and reports of this distinctive species from Arizona and New Mexico (A. J. Petrik-Ott 1979) have not been substantiated by herbarium specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cheilanthes yavapensis is an apogamous tetraploid, apparently formed by hybridization between C. lindheimeri and C. covillei (G. J. Gastony and M. D. Windham 1989). Although C. yavapensis has long been included within the concept of C. wootonii, the discovery that the similarities resulted from hybrid convergence rather than common ancestry requires that they be recognized as two distinct species. Unfortunately, the morphologic characteristics that separate these taxa are subtle, and careful study will be necessary to determine the proper dispositon of problematic specimens. In addition to the characteristics mentioned in the key, C. yavapensis is distinguished from C. wootonii by having larger spores, averaging more than 62 µm in diameter. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | Nephrodium lanosum, C. vestita | |
Name authority | (Michaux) D. C. Eaton: in Emory, Rep. U.S. Mex. Bound. 2(1): 234. (1859) | T. Reeves ex Windham: Contr. Univ. Michigan Herb. 19: 32. (1993) |
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