Cheilanthes lanosa |
Cheilanthes wootonii |
|
---|---|---|
hairy lip fern |
beaded lipfern, Wooton's lip fern |
|
Stems | compact to short-creeping, usually 4–8 mm diam.; scales often uniformly brown but at least some on each plant with thin, poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
long-creeping, 1–3 mm diam.; scales uniformly brown or weakly bicolored with poorly defined, dark, central stripe, lanceolate-ovate, straight to slightly contorted, loosely appressed, often deciduous on older portions of stem. |
Leaves | clustered, 7–50 cm; vernation circinate. |
scattered, 7–35 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
usually dark brown, rounded adaxially. |
Blade | linear-oblong to lanceolate, usually 2-pinnate-pinnatifid at base, 1.5–5 cm wide; rachis rounded adaxially, lacking scales, with monomorphic pubescence. |
oblong-lanceolate, 3–4-pinnate at base, 2–5 cm wide; rachis rounded adaxially, with scattered linear-lanceolate scales and sparse monomorphic pubescence. |
Ultimate segments | oblong to lanceolate, not beadlike, the largest 3–5 mm, abaxially and adaxially sparsely hirsute with long, segmented hairs. |
round to oblong, beadlike, the largest 1–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing sparsely hirsute adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | discontinuous, concentrated on small apical and lateral lobes. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 32 spores. |
Costae | brown adaxially for most of length; abaxial scales absent. |
green adaxially for most of length; abaxial scales multiseriate, lanceolate-ovate, truncate or subcordate at base, without overlapping basal lobes, conspicuous, the largest 0.4–0.8 mm wide, strongly imbricate, often concealing ultimate segments, ciliate, with coarse cilia often confined to proximal 1/2. |
n | = 2n = 90, apogamous. |
|
2n | = 60. |
|
Cheilanthes lanosa |
Cheilanthes wootonii |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, on a variety of substrates including limestone and granite | Rocky slopes and ledges, usually on igneous substrates |
Elevation | 100–800 m (300–2600 ft) | 800–2900 m (2600–9500 ft) |
Distribution |
AL; AR; CT; FL; GA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV |
AZ; CA; CO; NM; NV; OK; TX; UT; n Mexico
|
Discussion | Cheilanthes lanosa is apparently confined to the forests and prairies of eastern North America, and reports of this distinctive species from Arizona and New Mexico (A. J. Petrik-Ott 1979) have not been substantiated by herbarium specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Like its close relative Cheilanthes lindheimeri, C. wootonii is an apogamous triploid of unknown parentage. With the recognition of C. yavapensis as a distinct species, the name C. wootonii is restricted to populations with leaf blades that appear glabrous adaxially, costal scales that are often ciliate only in the proximal half, and stem scales that are usually brown and loosely appressed. In addition, C. wootonii is distinguished from C. yavapensis by having smaller spores, averaging less than 62 µm in diameter. These characteristics can be subtle, and some specimens will be difficult to place in either C. wootonii or C. yavapensis. T. Reeves (1979) identified several specimens from Arizona that he hypothesized were hybrids between C. wootonii and C. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | Nephrodium lanosum, C. vestita | |
Name authority | (Michaux) D. C. Eaton: in Emory, Rep. U.S. Mex. Bound. 2(1): 234. (1859) | Maxon: Proc. Biol. Soc. Wash. 3: 146. (1918) |
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