Cheilanthes lanosa |
Cheilanthes villosa |
|
---|---|---|
hairy lip fern |
villous lip fern |
|
Stems | compact to short-creeping, usually 4–8 mm diam.; scales often uniformly brown but at least some on each plant with thin, poorly defined, dark, central stripe, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
compact, 4–8 mm diam.; scales mostly bicolored, with broad, well-defined, dark, central stripe and narrow, light brown margins, linear-lanceolate, straight to slightly contorted, loosely appressed, persistent. |
Leaves | clustered, 7–50 cm; vernation circinate. |
clustered, 7–30 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
usually dark brown, rounded adaxially. |
Blade | linear-oblong to lanceolate, usually 2-pinnate-pinnatifid at base, 1.5–5 cm wide; rachis rounded adaxially, lacking scales, with monomorphic pubescence. |
oblong-lanceolate to ovate, 3–4-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, with scattered filiform to lanceolate scales, not pubescent. |
Ultimate segments | oblong to lanceolate, not beadlike, the largest 3–5 mm, abaxially and adaxially sparsely hirsute with long, segmented hairs. |
round to oval, beadlike, the largest 1–2 mm, abaxially nearly glabrous except for a few coarse hairs, adaxially villous with coarse, unbranched hairs. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing sparsely hirsute adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing villous adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal to obscurely inframarginal, slightly differentiated, 0.05–0.25 mm wide. |
Sori | discontinuous, concentrated on small apical and lateral lobes. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 32 spores. |
Costae | brown adaxially for most of length; abaxial scales absent. |
green adaxially for most of length; abaxial scales multiseriate, ovate to lanceolate, shallowly cordate at base, often with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, often concealing ultimate segments, erose-dentate, not ciliate. |
n | = 2n = 90, apogamous. |
|
2n | = 60. |
|
Cheilanthes lanosa |
Cheilanthes villosa |
|
Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes and ledges, on a variety of substrates including limestone and granite | Cliffs and rocky slopes, usually on limestone |
Elevation | 100–800 m (300–2600 ft) | 400–2200 m (1300–7200 ft) |
Distribution |
AL; AR; CT; FL; GA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV |
AZ; NM; TX; n Mexico |
Discussion | Cheilanthes lanosa is apparently confined to the forests and prairies of eastern North America, and reports of this distinctive species from Arizona and New Mexico (A. J. Petrik-Ott 1979) have not been substantiated by herbarium specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cheilanthes villosa is an apogamous triploid of unknown parentage. Although there are reports of hybridization between C. villosa and C. eatonii (D. B. Lellinger 1985), recent gene exchange is unlikely because both taxa are apogamous in North America. Morphologically intermediate specimens (tentatively called C. pinkavii in T. Reeves 1979) are included here in C. eatonii, pending further study and valid publication of Reeve's epithet. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | Nephrodium lanosum, C. vestita | |
Name authority | (Michaux) D. C. Eaton: in Emory, Rep. U.S. Mex. Bound. 2(1): 234. (1859) | Davenport ex Maxon: Proc. Biol. Soc. Wash. 31: 142. (1918) |
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